[1]Eulgem T, Rushton P, Robatzek S, Somssich IE. The WRKY superfamily of plant transcription factor. Trends Plant Sci, 2000, 5:199–206



[2]宋扬, 周军会, 张永强．植物组织特异性启动子研究.生物技术通报, 2007, 6:21–24



Song Y, Zhou J H, Zhang Y Q. Research on plant tissue-specific promoter. Biotechnol Bull, 2007, 6:21–24(in Chinese with English abstract)



[3]李志邈, 杨悦俭, 杨飞, 叶青静. 番茄根特异表达基因LeGRP2启动子的克隆及其在拟南芥的表达分析. 中国农业科学, 2010,43: 1877–1882



Li Z M, Yang Y J, Yang F, Ye Q J. Cloning of LeGRP2 promoter from tomato that shows root-specific expression in Arabidopsis. Sci Agric Sin,2010, 43: 1877–1882 (in Chinese with English abstract)



[4]杨学文, 任远, 田曾元, 张生学, 刘允军, 王国英.玉米茎特异启动子克隆及其在转基因烟草中的活性分析. 西北植物学报,2014,34:859–865



Yang X W, Ren Y, Tian Z Y, Zhang S X, Liu Y J, Wang G Y. Isolation of a shoot-specific promoter in maize and its activity in transgenic tobacco. Acta BotBor-OccidScin,2014,34:859–865(in Chinese with English abstract)



[5]颜彦, 林拥军. 一个水稻鞘叶特异表达启动子的克隆及功能鉴定. 农业生物技术学报, 2013, 21: 757–763



Yan Y, Lin Y J. Cloning and functional analysis of a promoter with tissue specific expression in rice (Oryza sativa Lssp. japonica) sheath and lea. JAgricBiotech, 2013, 21: 757–763(in Chinese with English abstract)



[6]Ulker B, Somssich I. WRKY transcription factors: from DNA binding towards biological function. CurrOpinPlant Biol, 2004, 7:491–498



[7]郝中娜, 王海华, 吴学龙, 郭泽建. 水稻WRKY19基因启动子功能研究初报. 中国农学通报, 2007, 23(6):99–103



Hao Z N, Wang H H, Wu X L, Guo Z J. Primary study of function of a rice WRKY19 gene promoter. Chin Agri Sci Bull, 2007, 23 (6):99–103 (in Chinese with English abstract)



[8]Paterson A H, Wendel J F, Gundlach H, Guo H, Jenkins J, Jin D,Llewellyn D, Showmaker K C, Shu S, Udall J, Yoo M, Byers R,Chen W, Doron-Faigenboim A, Duke M V, Gong L, Grimwood J,Grover C, Grupp K, Hu G, Lee T, Li J, Lin L, Liu T, Marler B S,Page J T, Roberts A W, Romanel E, Sanders W S, Szadkowski E,Tan X, Tang H, Xu C, Wang J, Wang Z, Zhang D, Zhang L,Ashfari H, Bedon F, Bowers J E, Brubaker C L, Chee P W, Das S,Gingle A R, Haigler C H, Harker D, Hoffmann L V, Hovav R,Jones D C, Lemke C, Mansoor S, Rahman M, Rainville L N,Rambani A, Reddy U K, Rong J, Saranga Y, Scheffler B E,Scheffler J A, Stelly D M, Triplett B A, Deynze A V, Vaslin M FS, Waghmare V N, Walford S, Wright R J, Zaki E A, Zhang T,Dennis E S, Mayer K F X, Peterson D G, Rokhsar D S, Wang X,Schmutz J. Repeated polyploidization of Gossypium genomes and the evolution of spinnable cotton fibres. Nature, 2012, 492:423–427



[9]Wang K B, Wang Z W, Li F G, Ye W W, Wang J Y, Song G L, Yue Z, Cong L, Shang H H, Zhu S L, Zou C S, LI Q, Yuan Y L, Lu C R, Wei H L, Gou C Y, Zheng Z Q, Yin Y, Zhang X Y, Liu K, Wang B, Song C, Shi N, Kohel R J, Percy R G, Yu J Z, Zhu Y X, Wang J, Yu S X. The draft genome of a diploid cotton Gossypiumraimondii. Nat Genet, 2012, 44: 1098–1103



[10]Cai C P,Niu E L, Du H,Zhao L,Feng Y,Guo W Z. Genome-wide analysis of the WRKY transcriptionfactor gene family in Gossypium raimondii and theexpression of orthologs in cultivated tetraploid cotton.Crop J,2014,2:87–101



[11]Paterson A H, Brubaker C, Wendel J F. A rapid method for extraction of cotton (Gossypium spp.) genomic DNA suitable for RFLP or PCR analysis. Plant Mol Biol Rep, 1993, 11: 122–127



[12]马雪梅, 胥晓, 李晓波, 肖向文. 农杆菌介导快速、高效获得转基因烟草纯合株系. 中国烟草学报, 2012,18(4):66–71



Ma X M, Xu X, Li X B, Xiao X W. Rapid and efficient production of homozygous transgenic tobacco plants Agrobacterium tumefaciens.Acta TobSci,2012,18(4):66–71 (in Chinese with English abstract)



[13]Livak K J, Schmittgen T D. Analysis of relative gene expression data using real-time quantitative PCR and the 2–ΔΔCT method. Methods, 2001, 25: 402–408



[14]Jefferson R A. Assaying chimeric genes in plants: The GUS gene fusion system. Plant MolBiolRep, 1987, 5: 387–405



[14]王雪,谭艳平, 周杰, 王春台, 刘学群.一个烟草糖基转移酶启动子在转基因烟草植株中的表达分析.安徽农业科学, 2010, 38:9426–9428



Wang X, Tang Y P, Zhou J, Wang C T, Liu X Q.Expression of a tobacco Glycosyl transferasegene’s promoter in transgenic tobacco. JAnhui AgriSci,2010,38:9426–9428(in Chinese with English abstract)



[15]付永平, 周海涛, 王丕武.大豆种子特异性启动子的克隆及功能分析.西北农林科技大学学报(自然科学版), 2009, 37(12): 105–111



Fu Y P, Zhou H T, Wang P W.Cloning and identification of the seed specific promoter from soybean.JNorthwest SciTech UnivAgricFor(NatSciEdn), 2009, 37(12):105–111 (in Chinese with English abstract)



[16]崔同霞, 白江平, 魏桂民, 赵旭, 王蒂, 张金文. 马铃薯SGT3基因表达及其启动子功能分析. 草业学报, 2014, 23(2): 196–206



Cui T X, Bai J P, Wei G M, Zhao X, Wang D, Zhang J W.Gene expression and function analysis of SGT3 promoter from Potato. Acta PraSin,2014,23(2): 196–206(in Chinese with English abstract)



[17]许兰珍, 彭爱红, 何永睿, 姚利晓, 雷天刚, 刘小丰, 姜国金, 邹修平, 陈善春. 异源韧皮部特异启动子在转基因枳中的表达.园艺学报, 2014, 41: 1–8



Xu L Z, Peng A H, He Y R, Yao L X, Lei T G, Liu X F, Jiang G J, Zou X P, Chen S C.Expression analysis of three phloem-specific promoters in transgenic Poncirus trifoliate. Acta HorSin, 2014, 41:1–8(in Chinese with English abstract)



[18]房孝良, 刘炜, 安静,王庆国. 水稻种胚特异性启动子OsESP1的克隆及其表达特性. 作物学报, 2011, 37: 1904−1909



Fang X L, Liu W, An J, Wang Q G. Isolation and characterization of an embryo-specific promoter OsESP1 from Rice. Acta AgronSin, 2011, 37: 1904−1909(in Chinese with English abstract)



[19]Antonio A,Rodrigo R,Luciane M,Naiara P,Osmundo B,Sinara A,SarahM N,Marcio A,João A,Maria C, Maria F. Isolation and functional characterization of a cotton ubiquitination-related promoter and 5?UTR that drives high levels of expression in root and flower tissues.BMC Biotechnol, 2011, 11:115



[20]Lightfoot D J, Orford S J, Timmis J N. Identification and characterisation of cotton boll wall-specific gene promoters for future transgenic cotton varieties. Plant MolBiolRep,2013,31: 174–184



[21]Zahur M,Maqbool A, Irfan M,Younas M, Barozai K,Qaiser U,Rashid B , Husnain T, Riazuddin S. Functional analysis of cotton small heat shock protein promoter region in response to abiotic stresses in tobacco using Agrobacterium-mediated transient assay. MolBiolRep, 2009, 36:1915–1921



[22]Wu A M, Hu J S., Liu J Y. Functional analysis of a cotton cellulose synthase A4 gene promoter in transgenic tobacco plants. Plant Cell Rep, 2009, 28:1539–1548



[23]RyuHS,HanM,LeeSK,ChoJI,RyooN,HeuS,LeeYH,BhooSH,WangGL,HahnTR,JeonJS.AcomprehensiveexpressionanalysisoftheWRKYgenesuperfamilyinriceplantsduringdefenseresponse.Plant Cell Rep, 2006, 25:836–847



[24]Wu X L, Shiroto Y, Kishitani S, Ito Y, Toriyama K.Enhanced heat and drought tolerance in transgenic rice seedlings over-expressing OsWRKY11 under the control of HSP101 promoter.Plant Cell, 2009, 28:21–30



[25]Qiu Y P,Yu D Q.Over-expression of the stress induced OsWRKY45 enhances disease resistance and drought tolerance in Arabidopsis. EnvironExpBot,2009,65:35–47



[26]付乾堂, 余迪求. 拟南芥AtWRKY25,AtWRKY26和AtWRKY33在非生物胁迫条件下的表达分析.遗传, 2010, 32:848–856



Fu Q T, Yu D Q.Expression profiles of AtWRKY25, AtWRKY26 and AtWRKY33 under abiotic stresses. Hereditas(Beijing),2010, 32:848–856(in Chinese with English abstract)



[27]王磊, 高晓清, 朱苓华, 周永力, 黎志康.植物WRKY转录因子家族基因抗病相关功能的研究进展. 植物遗传资源学报, 2011, 12: 80–85



Wang L, Gao X Q, Zhu L H, Zhou Y L, Li Z K. Advances in research on function of WRKY transcription factor genes in plant resistance.JPlant GenRes, 2011, 12: 80–85(in Chinese with English abstract)



[28]Wu K L, Guo Z J, Wang H H, Li J. The WRKY family of transcription factors in rice and Arabidopsis and their origins. DNA Res, 2005, 12: 9–26



[29]范荣伟, 谢华, 姚磊, 马荣才.大白菜BrWRKY33基因上游调控序列的克隆及功能分析.农业生物技术学报, 2010, 18:670–675



Fan R W, Xie H, Yao L, Ma R C. Cloning and expression analysis of the up-stream regulatory sequence of transcription factor gene BrWRKY33 from Chinese cabbage. JAgricBiotech,2010, 18:670–675(in Chinese with English abstract)



[30]张春晓, 王文棋, 蒋湘宁, 陈雪梅.植物基因启动子研究进展.遗传学报, 2004,31: 1455–1464



Zhang C X, Wang W Q, Jiang X N, Chen X M.Review on plant gene promoter. Acta GenetSin, 2004,31:1455–1464(in Chinese with English abstract)