[1]Kang G Z, Peng X Q, Wang L, Yang Y Y, Shao R X, Xie Y X, Ma D Y, Wang C Y, Guo T C, Zhu Y J. Ultrastructural observation of mesophyll cells and temporal expression profiles of the genes involved in transitory starch metabolism in flag leaves of wheat after anthesis. Physiol Plant, 2015, 153: 12–29
[2]Xue Q W, Zhu Z X, Musick J T, Stewart B A, Dusek D A. Physiological mechanisms contributing to the increased water-use efficiency in winter wheat under deficit irrigation. J Plant Physiol, 2006, 163: 154–164
[3]Kohl S, Hollmann J, Erban A, Kopka J, Riewe D, Weschke W, Weber H. Metabolic and transcriptional transitions in barley glumes reveal a role as transitory resource buffers during endosperm filling. J Lipid Res, 2008, 49: 880–892
[4]张永平, 王志敏, 王璞, 赵明. 冬小麦节水高产栽培群体光合特征. 中国农业科学, 2003, 36: 1143–1149
Zhang Y P, Wang Z M, Wang P, Zhao M. Canopy photosynthetic characteristics of population of winter wheat in water-saving and high-yielding cultivation. Sci Agric Sin, 2003, 36: 1143–1149 (in Chinese with English abstract)
[5]Gebbing T, Schnyder H. 13C Labeling kinetics of sucrose in glumes indicates significant refixation of respiratory CO2 in the wheat ear. Funct Plant Biol, 2001, 28: 1047–1053
[6]王志敏, 张英华, 张永平,吴永成. 麦类作物穗器官的光合性能研究进展. 麦类作物学报, 2004, 24(4): 136–139
Wang Z M, Zhang Y H, Zhang Y P, Wu Y C. Review on photosynthetic performance of ear organs in Triticeae crops. J Triticeae Crops, 2004, 24(4): 136–139 (in Chinese with English abstract)
[7]张英华, 苏达, 张胜全, 周顺利, 王志敏, 张永平, 方保停. 不同水分条件下冬小麦旗叶和穗器官的PEPC活性及其与粒重和蛋白质含量的关系. 麦类作物学报, 2009, 29: 997–1003
Zhang Y H, Su D, Zhang S Q, Zhou S L, Wang Z M, Zhang Y P, Fang B T. Phosphoenolpyruvate carboxylase activity of flag leaf and ear organs and its relationship with grain mass and protein content in winter wheat under different water treatments. J Triticeae Crops, 2009, 29: 997–1003 (in Chinese with English abstract)
[8]李朝霞, 赵世杰, 孟庆伟, 邹琦, 田纪春. 不同粒叶比小麦品种非叶片光合器官光合特性的研究. 作物学报, 2004, 30: 419–426
Li Z X, Zhao S J, Meng Q W, Zou Q, Tian J C. Photosynthetic characteristics in non-leaf organs of winter wheat cultivars differing in grain-leaf ratio. Acta Agron Sin, 2004, 30: 419–426 (in Chinese with English abstract)
[9]Tambussi E A, Bort J, Guiamet J J. The photosynthetic role of ears in C3 cereals: metabolism, water use efficiency and contribution to grain yield. Crit Rev Plant Sci, 2007, 26: 1–16
[10]Sanchez-Bragado R, Elazab A, Zhou B W, Serret M D, Bort J, Nieto-Taladriz M T, Araus J L. Contribution of the ear and the flag leaf to grain filling in durum wheat inferred from the carbon isotope signature: genotypic and growing conditions effects. J Integr Plant Biol, 2014, 56: 444–454
[11]王维, 张建华, 杨建昌,朱庆森. 适度土壤干旱对贪青小麦茎鞘贮藏性糖运转及籽粒充实的影响. 作物学报, 2004, 30: 1019–1025
Wang W, Zhang J H, Yang J C, Zhu Q S. Effects of controlled soil drought on remobilization of stem-stored carbohydrate and grain filling of wheat with unfavorably-delayed senescence. Acta Agron Sin, 2004, 30: 1019–1025 (in Chinese with English abstract)
[12]Araus J L, Brown H R, Febrero A, Bort J, Serret M D. Ear photosynthesis, carbon isotope discrimination and the contribution of respiratory CO2 to differences in grain mass in durum wheat. Plant Cell & Environ, 1993, 16: 383–392
[13]Maydup M L, Antonietta M, Guiamet J J, Graciano C, López J.R, Tambussi E A. The contribution of ear photosynthesis to grain filling in bread wheat (Triticum aestivum L.). Field Crops Res, 2010, 119: 48–58
[14]Aranjuelo I, Cabrerabosquet L, Morcuende R, Avice J C, Nogués S, Araus J L, Martínez-Carrasco R, Pérez P. Does ear C sink strength contribute to overcoming photosynthetic acclimation of wheat plants exposed to elevated CO2? J Exp Bot, 2011, 62: 3957–3969
[15]张磊, 吕金印, 贾少磊. 水分亏缺对小麦穗部光合特性及花前14C-同化物分配的影响. 作物学报, 2013, 39: 1514–1519
Zhang L, Lyu J Y, Jia S L. Photosynthetic characteristics of spike and distribution of 14C-assimilates accumulated before anthesis in wheat under water deficit condition. Acta Agron Sin, 2013, 39: 1514–1519 (in Chinese with English abstract)
[16]任妍婷, 吕金印, 成健. 水分亏缺对不同抗旱性小麦穗部光合及碳同化物转运的影响. 麦类作物学报, 2012, 32: 683–688
Ren Y T, Lyu J Y, Cheng J. Effects of water deficit on photosynthetic characteristics, accumulation and transportation of 14C-assimilates of ears in wheat. J Triticeae Crops, 2012, 32: 683–688 (in Chinese with English abstract)
[17]Jia S L, Lyu J L, Jiang S X, Liu C X, Jing Z H. Response of wheat ear photosynthesis and photosynthate carbon distribution to water deficit. Photosynthetica, 2015, 53: 95–109
[18]Teare I D, Peterson C J. Surface area of chlorophyll-containing tissue on the inflorescence of Triticum aestivum L. Crop Sci, 1971,11: 627–628
[19]裘昭峰, 翟立业. 小麦穗和芒表面积的估测. 作物学报, 1985, 11: 138, 144
Qiu S F, Zhai L Y. The estimation for surface area of spike and awn of the common wheat. Acta Agron Sin, 1985, 11: 138, 144 (in Chinese)
[20]Martinez D E, Luquez V M, Bartoli C G, Guiamét J J. Persistence of photosynthetic components and photochemical efficiency in ears of water-stressed wheat (Triticum aestivum). Physiol Plant, 2003, 119: 519–525
[21]魏爱丽, 王志敏, 陈斌, 翟志席, 张英华. 土壤干旱对小麦绿色器官光合电子传递和光合磷酸化活力的影响. 作物学报, 2004, 30: 487–490
Wei A L, Wang Z M, Chen B, Zhai Z X, Zhang Y H. Effect of soil drought on electron transport rate and photophosphorylation level of different green organs in wheat. Acta Agron Sin, 2004, 30: 487–490 (in Chinese with English abstract)
[22]Gallagher J N, Biscoe P V, Hunter B. Effects of drought on grain growth. Nature, 1976, 264: 541–542
[23]Yang J C, Zhang J H, Huang Z L, Zhu Q S, Wang L. Remobilization of carbon reserves is improved by controlled soil-drying during grain filling of wheat. Crop Sci, 2000, 40: 1645–1655
[24]王维, 蔡一霞, 张建华,杨建昌, 朱庆森. 适度土壤干旱对贪青小麦茎贮藏碳水化合物向籽粒运转的调节. 作物学报, 2005, 31: 289–296
Wang W, Cai Y X, Zhang J H, Yang J C, Zhu Q S. Regulation of controlled soil drying on remobilization of stem-stored carbohydrate to grain in wheat grown under unfavorably-delayed senescence. Acta Agron Sin, 2005, 31: 289–296 (in Chinese with English abstract)
[25]Tambussi E, Nogues S J. Ear of durum wheat under water stress: water relations and photosynthetic metabolism. Planta, 2005, 221: 446–458
[26]Bidinger F, Musgrave R B, Fischer R A. Contribution of stored pre-anthesis assimilate to grain yield in wheat and barley. Nature, 1977, 270: 431–433
[27]Evans L T, Bingham J, Jackson P, Sutherland J. Effect of awns and drought on the supply of photosynthate and its distribution within wheat ears. Ann Appl Biol, 1972, 70: 67–76
[28]Abebe T, Wise R P, Skadsen R W. Comparative transcriptional profiling established the awn as the major photosynthetic organ of the barley spike while the lemma and the palea primarily protect the seed. Plant Genome, 2009, 2: 247–259
[29]Ziegler-Jöns A. Gas exchange of ears of cereals in response to carbon dioxide and light: I. Relative contributions of parts of the ears of wheat, oat, and barley to the gas exchange of the whole organ. Planta, 1989, 178: 84–91
[30]Olugbemi L B. Distribution of carbon-14 assimilated by wheat awns. Ann Appl Biol, 1978, 90: 111–114