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作物学报 ›› 2020, Vol. 46 ›› Issue (4): 571-585.doi: 10.3724/SP.J.1006.2020.92044

• 耕作栽培·生理生化 • 上一篇    下一篇

安徽沿淮地区优质高产常规粳稻品种筛选及特征特性

卫平洋1,裘实2,唐健1,肖丹丹1,朱盈1,刘国栋1,邢志鹏1,胡雅杰1,郭保卫1,高尚勤3,魏海燕1,*(),张洪程1,*()   

  1. 1 扬州大学江苏省作物遗传生理重点实验室 / 扬州大学农业部长江流域稻作技术创新中心 / 粮食作物现代产业技术协同创新中心, 江苏扬州 225009
    2 江苏省农业科学院种质资源与生物技术研究所, 江苏南京 210014
    3 安徽省凤台县农业技术推广中心, 安徽淮南 232100
  • 收稿日期:2019-08-14 接受日期:2019-12-26 出版日期:2020-04-12 网络出版日期:2020-01-15
  • 通讯作者: 魏海燕,张洪程
  • 作者简介:E-mail: 2918052418@qq.com
  • 基金资助:
    本研究由国家重点研发计划项目(2016YFD0300503);江苏省重点研发计划项目(BE2016344);江苏省农业科技自主创新基金(CX[15]1002);扬州大学拔尖人才计划和江苏省农业三新工程资助(SXGC[2017]294)

Screening and characterization of high-quality and high-yield japonica rice varieties in Yanhuai region of Anhui province

WEI Ping-Yang1,QIU Shi2,TANG Jian1,XIAO Dan-Dan1,ZHU Ying1,LIU Guo-Dong1,XING Zhi-Peng1,HU Ya-Jie1,GUO Bao-Wei1,GAO Shang-Qin3,WEI Hai-Yan1,*(),ZHANG Hong-Cheng1,*()   

  1. 1 Jiangsu Key Laboratory of Crop Genetics and Physiology / Innovation Center of Rice Cultivation Technology in Yangtze Valley, Ministry of Agriculture / Co-Innovation Center for Modern Production Technology of Grain Crops, Yangzhou University, Yangzhou 225009, Jiangsu, China
    2 Institute of Germplasm Resources and Biotechnology, Jiangsu Academy of Agricultural Sciences, Nanjing 210014, Jiangsu, China
    3 Extension Center of Agriculture Technology in Fengtai County of Anhui Province, Huainan 232100, Anhui, China
  • Received:2019-08-14 Accepted:2019-12-26 Published:2020-04-12 Published online:2020-01-15
  • Contact: Hai-Yan WEI,Hong-Cheng ZHANG
  • Supported by:
    This study was supported by the National Key Research Program(2016YFD0300503);the Key Research Program of Jiangsu Province(BE2016344);the Major Independent Innovation Project in Jiangsu Province(CX[15]1002);the Program for Scientific Elitists of Yangzhou University, and the Three New Agricultural Engineering Fund of Jiangsu Province(SXGC[2017]294)

摘要:

以74个常规粳稻品种(品系)为材料, 根据产量和食味值分为高产优质、高产不优质、中产优质和中产不优质4种类型。选用高产优质、高产不优质和中产优质3种类型共9个品种, 研究其产量、品质差异及其高产优质形成的特征特性, 以期为安徽沿淮地区适宜品种的筛选、种植推广提供科学依据和理论支撑。结果表明, 中产类型与高产类型粳稻在产量上有极显著差异, 2017年和2018年, 中产优质类型产量比高产优质类型、高产不优质类型分别低16.95%、16.76%和16.52%、16.33%。2017年, 与高产不优质类型相比, 高产优质类型和中产优质类型的直链淀粉含量分别低39.31%、42.63%; 胶稠度分别长22.06%、19.12%; 蛋白质含量分别低11.60%、17.78%。这些高产优质粳稻品种特征特性主要表现为, 产量在8.35~9.16 t hm -2, 单位面积穗数在310×10 4~320×10 4 hm -2之间, 每穗粒数在140左右, 千粒重在25 g以上; 食味值评分在60~74, 胶稠度长度在80~90 mm, 蛋白质含量在6%~8%。

关键词: 沿淮地区, 粳稻, 产量, 品质, 品种筛选

Abstract:

A field experiment was conducted with 74 conventional japonica rice varieties (lines), which were divided into four types according to the yield and taste value, i.e. high yield with good quality (HG), high yield with bad quality (HB), medium yield with good quality (MG), and medium yield with bad quality (MB). Among them, HG, HB, MG were weed to analyze their yield, quality differences and characteristics in HG formation, providing a scientific basis and theoretical support for screening, planting and extending the suitable varieties in Yanhuai area of Anhui province. The yield of MG was 16.95%, 16.76%, and 16.52%, 16.33% lower than that of HG type and HB type, respectively in 2017 and 2018. In 2017, compared with HB type, HG and MG types decreased by 39.31% and 42.63% in amylose content, increased by 22.06% and 19.12% in length of gel consistency, and decreased by 11.60% and 17.78% in protein content. The characteristics of these HG japonica rice varieties were mainly as follows: the yield was 8.35-9.16 t hm -2, the number of panicles per unit area was 310×10 4-320×10 4 hm -2, the grain number per panicle was around 140, the 1000-grain weight was over 25 g; the taste value was 60-74, the length of gel consistency was 80-90 mm, and the protein content was between 6% and 8%.

Key words: Yanhuai region, japonica rice, yield, quality, variety screening

表1

常规粳稻产量及其食味值"

年份
Year
指标
Index
变幅大小
Variable amplitude
均值
Mean
标准差
Standard deviation
变异系数
CV(%)
2017 食味值 Taste value 40-71 55 7.95 14.37
产量 Yield (t hm-2) 5.69-9.28 7.67 0.77 10.05
2018 食味值 Taste value 43-74 57 7.84 12.48
产量 Yield (t hm-2) 5.51-9.21 7.53 0.82 11.50

表2

不同类型常规粳稻产量、食味值和生育期的差异"

品种
类型
Type
品种
数量
Number
品种名称
Cultivar
食味值Taste value 产量Yield (t hm-2) 生育期
Growth period (d)
均值
Mean
变幅
Variable amplitude
均值
Mean
变幅
Variable amplitude
高产
优质
HG
16 南粳9108, 南粳505, 南粳5718, 宁9039, 武4610, 常软07-6, 常软07-1, 南粳2728, 镇稻99, 徐稻9号, 宁9003, 宁9022, 常软07-4, 常软07-3, 宁5720, 常软07-2
Nanjing 9108, Nanjing 505, Nanjing 5718, Ning 9039, Wu 4610, Changruan 07-6, Changruan 07-1, Nanjing 2728, Zhendao 99, Xudao 9, Ning 9003, Ning 9022, Changruan 07-4, Changruan 07-3, Ning 5720, Changruan 07-2
65 60-74 8.68 8.35-9.16 150-160
高产
不优质
HB
15 武运5051, 扬育粳2号, 扬粳1612, 华粳8号, 连粳12, 宁粳040, 泗15-301, 华粳5号, 武运粳21, 镇9471, 连粳13, 连粳13264, 苏秀867, 连粳15, 武运5020
Wuyun 5051, Yangyujing 2, Yangjing 1612, Huajing 8, Lianjing 12, Ningjing 040, Si 15-301, Huajing 5, Wuyunjing 21, Zhen 9471, Lianjing 13, Lianjing 13264, Suxiu 867, Lianjing 15, Wuyun 5020
53 45-59 8.60 8.24-9.28 140-160
中产
优质
MG
11 宁粳7号, 宁粳4号, 淮稻5号, 常软07-5, 福粳1606, 沪香粳165, 南繁1610, 南繁1605, 泗15-234, 苏香粳3号, 松早香1号
Ningjing 7, Ningjing 4, Huaidao 5, Changruan 07-5, Fujing 1606, Huxiangjing 165, Nanfan 1610, Nanfan 1605, Si 15-234, Suxiangjing 3, Songzaoxiang 1
65 63-69 7.57 6.88-8.05 135-155
中产
不优质
MB
32 盐粳16, 丰粳3227, 新稻22, 圣稻18-15, 徐36618, 苏粳815, 连粳11, 泗稻14-211, 淮330, 南粳5711, 泰粳2340, 武运粳80, 华丰1502, 泰粳1152, 华粳295, 徐农33202, 南粳5833, 扬粳5515, 扬粳3491, 扬粳3012, 武运粳32, 武运粳27, 泗276, 泗稻15, 皖垦粳3号, 武粳004, 圣稻1647, 圣稻19, 圣稻20, 新科稻31, 日光, 连粳15113
Yanjing 16, Fengjing 3227, Xindao 22, Shengdao 18-15, Xu 36618, Sujing 815, Lianjing 11, Sidao 14-211, Huai 330, Nanjing 5711, Taijing 2340, Wuyunjing 80, Huafeng 1502, Taijing 1152, Huajing 295, Xunong 33202, Nanjing 5833, Yangjing 5515, Yangjing 3491, Yangjing 3012, Wuyunjing 32, Wuyunjing 27, Si 276, Sidao 15, Wankenjing 3, Wujing 004, Shengdao 1647, Shengdao 19, Shengdao 20, Xinkedao 31, Riguang, Lianjing 15113
50 40-59 7.25 5.51-7.96 135-155

表3

不同类型常规粳稻的产量及其构成因素"

品种类型
Type
品种名称
Cultivar
单位面积穗数
No. of panicles
(×104 hm-2)
每穗粒数Spikelets per panicle 结实率Seed-setting
rate (%)
千粒重1000-grain
weight (g)
产量
Yield
(t hm-2)
2017
高产优质
HG
南粳9108 Nanjing 9108 318.20 139.98 86.80 25.13 9.08
南粳505 Nanjing 505 314.80 139.49 84.61 25.28 8.89
徐稻9号Xudao 9 316.40 140.32 83.69 25.40 8.93
平均值Mean 316.47 Ab 139.93 Aa 85.03 Aa 25.27 Aa 8.97 Aa
高产不优质
HB
武运5051 Wuyun 5051 320.40 140.33 84.91 24.58 8.80
扬粳1612 Yangjing 1612 323.50 138.26 85.67 24.30 8.99
华粳8号Huajing 8 324.10 136.34 83.81 25.28 9.05
平均值Mean 322.67 Ab 138.31 Aa 84.80 Aa 24.72 Aab 8.95 Aa
中产优质
MG
福粳1606 Fujing 1606 345.00 112.80 82.47 23.50 7.54
苏香粳3号Suxiangjing 3 396.55 91.50 82.39 23.53 7.03
松早香1号 Songzaoxiang 1 346.80 109.50 82.38 24.93 7.79
平均值Mean 362.78 Aa 104.60 Bb 82.41 Ab 23.99 Ab 7.45 Bb
2018
高产优质
HG
南粳9108 Nanjing 9108 319.20 138.78 86.20 25.85 9.15
南粳505 Nanjing 505 310.75 137.00 85.53 25.90 8.98
徐稻9号Xudao 9 311.45 138.02 85.46 25.25 8.93
平均值Mean 313.80 Ab 137.93 Aa 85.73 Aa 25.67 Aa 9.02 Aa
高产不优质
HB
武运5051 Wuyun 5051 315.50 137.17 85.30 24.83 8.92
扬粳1612 Yangjing 1612 319.20 137.95 84.82 25.35 9.02
华粳8号Huajing 8 316.45 136.44 85.90 25.55 9.05
平均值Mean 317.05 Ab 137.19 Aa 85.34 Aa 25.24 Aab 9.00 Aa
中产优质
MG
福粳1606 Fujing 1606 348.70 113.46 81.96 23.90 7.72
苏香粳3号Suxiangjing 3 385.70 94.00 83.68 23.73 7.22
松早香1号 Songzaoxiang 1 345.25 107.00 83.46 24.80 7.64
平均值Mean 359.88 Aa 104.82 Bb 83.03 Ab 24.14 Ab 7.53 Bb

表4

不同类型常规粳稻的食味值"

品种类型
Type
品种名称
Cultivar
食味值
Taste value
外观
Appearance
硬度
Hardness
黏度
Viscosity
平衡度
Balance degree
2017
高产优质
HG
南粳9108 Nanjing 9108 71 6.8 6.5 6.9 6.7
南粳505 Nanjing 505 69 6.7 6.7 6.5 6.4
徐稻9号Xudao 9 71 6.7 6.6 7.1 6.8
平均值Mean 70 Aa 6.7 Aa 6.6 Bb 6.8 Aa 6.6 Aa
高产不优质
HB
武运5051 Wuyun 5051 58 5.3 7.2 4.5 4.7
扬粳1612 Yangjing 1612 60 4.9 7.5 5.5 4.9
华粳8号Huajing 8 59 4.9 7.5 5.4 4.9
平均值Mean 59 Bb 5.0 Bb 7.4 Aa 5.1 Ab 4.8 Bb
中产优质
MG
福粳1606 Fujing 1606 69 6.8 6.6 6.4 6.4
苏香粳3号Suxiangjing 3 71 7.0 6.5 7.3 7.1
松早香1号 Songzaoxiang 1 67 6.5 6.6 5.9 6.1
平均值Mean 69 Aa 6.8 Aa 6.6 Bb 6.5 Aa 6.5 Aa
2018
高产优质
HG
南粳9108 Nanjing 9108 68 6.3 6.8 6.5 6.3
南粳505 Nanjing 505 73 7.0 6.3 6.9 7.4
徐稻9号Xudao 9 71 6.5 6.5 6.8 7.2
平均值Mean 71 Aa 6.6 Aa 6.5 Bb 6.7 Aa 7.0 Aa
高产不优质
HB
武运5051 Wuyun 5051 57 5.5 7.5 4.8 4.7
扬粳1612 Yangjing 1612 59 5.4 7.3 5.8 4.5
华粳8号Huajing 8 57 4.9 7.5 5.4 4.5
平均值Mean 58 Bb 5.3 Bb 7.4 Aa 5.3 Ab 4.6 Bb
中产优质
MG
福粳1606 Fujing 1606 68 6.3 6.9 6.6 6.2
苏香粳3号Suxiangjing 3 69 6.9 6.7 7.2 6.9
松早香1号 Songzaoxiang 1 71 6.7 6.7 5.9 6.8
平均值Mean 69 Aa 6.6 Aa 6.8 Bb 6.6 Aa 6.6 Aa
年度间 Between years ns ns ns ns ns

表5

不同类型常规粳稻的加工品质"

品种类型
Type
品种名称
Cultivar
糙米率
Brown rice rate
精米率
Milled rice rate
整精米率
Head milled rice rate
2017
高产优质
HG
南粳9108 Nanjing 9108 85.58 69.56 64.29
南粳505 Nanjing 505 82.86 66.19 62.09
徐稻9号Xudao 9 84.28 71.62 59.54
平均值Mean 84.24 Aa 69.12 Aa 61.97 Aab
高产不优质
HB
武运5051 Wuyun 5051 82.24 64.69 63.19
扬粳1612 Yangjing 1612 83.47 68.06 61.76
华粳8号Huajing 8 83.33 72.09 64.38
平均值Mean 83.01 Aa 68.28 Aa 63.11 Aa
中产优质
MG
福粳1606 Fujing 1606 83.25 69.58 58.80
苏香粳3号Suxiangjing 3 84.68 69.39 60.03
松早香1号 Songzaoxiang 1 84.03 70.58 57.12
平均值Mean 83.99 Aa 69.85 Aa 58.65 Ab
2018
高产优质
HG
南粳9108 Nanjing 9108 84.93 70.94 64.06
南粳505 Nanjing 505 83.22 66.70 62.37
徐稻9号Xudao 9 84.28 69.75 60.13
平均值Mean 84.14 Aa 69.13 Aa 62.19 Aab
高产不优质
HB
武运5051 Wuyun 5051 82.68 66.63 62.67
扬粳1612 Yangjing 1612 83.64 66.88 62.01
华粳8号Huajing 8 83.50 70.30 63.53
平均值Mean 83.27 Aa 67.94 Aa 62.74 Aa
中产优质
MG
福粳1606 Fujing 1606 84.66 70.65 58.98
苏香粳3号Suxiangjing 3 83.56 70.53 60.27
松早香1号 Songzaoxiang 1 83.50 71.04 57.96
平均值Mean 83.91 Aa 70.74 Aa 59.07 Ab
年度间 Between years ns ns ns

表6

不同类型常规粳稻的外观品质"

品种类型
Type
品种名称
Cultivar
透明度
Transparency
长宽比Length-width ratio 垩白率
Chalkiness rate (%)
垩白度
Chalkiness degree (%)
2017
高产优质
HG
南粳9108 Nanjing 9108 3 1.67 64.95 23.30
南粳505 Nanjing 505 4 1.75 59.82 21.86
徐稻9号Xudao 9 2 1.83 42.57 19.57
平均值Mean 3 Aa 1.75 Aa 55.78 Aa 21.58 Aa
高产不优质
HB
武运5051 Wuyun 5051 3 1.76 49.46 18.73
扬粳1612 Yangjing 1612 2 1.75 47.95 16.24
华粳8号Huajing 8 2 1.93 22.73 8.15
平均值Mean 2 Aa 1.81 Aa 40.05 Ab 14.37 Ab
中产优质
MG
福粳1606 Fujing 1606 3 1.70 45.02 18.68
苏香粳3号Suxiangjing 3 4 2.03 66.85 23.75
松早香1号 Songzaoxiang 1 3 2.35 49.31 18.77
平均值Mean 3 Aa 2.03 Aa 53.73 Aa 20.40 Aab
2018
高产优质
HG
南粳9108 Nanjing 9108 3 1.71 67.05 25.80
南粳505 Nanjing 505 5 1.74 61.86 22.85
徐稻9号Xudao 9 3 1.89 42.83 20.34
平均值Mean 4 Aa 1.78 Aa 57.25 Aa 23.00 Aa
高产不优质
HB
武运5051 Wuyun 5051 4 1.76 46.63 18.87
扬粳1612 Yangjing 1612 3 1.73 46.28 19.79
华粳8号Huajing 8 2 1.92 21.57 8.43
平均值Mean 3 Aa 1.80 Aa 38.16 Ab 15.70 Ab
中产优质
MG
福粳1606 Fujing 1606 2 1.76 49.92 20.47
苏香粳3号Suxiangjing 3 4 2.11 66.21 25.85
松早香1号 Songzaoxiang 1 4 2.32 48.73 19.26
平均值Mean 3 Aa 2.06 Aa 54.95 Aa 21.86 Aab
年度间 Between years ns ns ns ns

表7

不同类型常规粳稻的营养品质及蒸煮食味品质"

品种类型
Type
品种名称
Cultivar
直链淀粉含量
Amylose content (%)
胶稠度
Gel consistency (mm)
蛋白质含量
Protein content (%)
2017
高产优质
HG
南粳9108 Nanjing 9108 10.49 87 7.59
南粳505 Nanjing 505 10.43 82 7.05
品种类型
Type
品种名称
Cultivar
直链淀粉含量
Amylose content (%)
胶稠度
Gel consistency (mm)
蛋白质含量
Protein content (%)
徐稻9号Xudao 9 13.87 81 7.40
平均值Mean 11.60 Ab 83 Aa 7.35 Ab
高产不优质
HB
武运5051 Wuyun 5051 14.17 67 7.99
扬粳1612 Yangjing 1612 16.32 71 7.64
华粳8号Huajing 8 18.00 66 8.16
平均值Mean 16.16 Aa 68 Bb 7.93 Aa
中产优质
MG
福粳1606 Fujing 1606 11.51 81 7.55
苏香粳3号Suxiangjing 3 11.75 78 7.69
松早香1号 Songzaoxiang 1 10.72 83 7.21
平均值Mean 11.33 Ab 81 Aa 7.55 Aab
2018
高产优质
HG
南粳9108 Nanjing 9108 10.47 89 7.43
南粳505 Nanjing 505 10.41 83 7.04
徐稻9号Xudao 9 13.67 81 7.34
平均值Mean 11.52 Ab 84 Aa 7.27 Ab
高产不优质
HB
武运5051 Wuyun 5051 13.91 65 7.90
扬粳1612 Yangjing 1612 16.42 68 7.59
华粳8号Huajing 8 17.84 65 8.10
平均值Mean 16.06 Aa 66 Bb 7.86 Aa
中产优质
MG
福粳1606 Fujing 1606 11.20 80 7.69
苏香粳3号Suxiangjing 3 11.51 78 7.83
松早香1号 Songzaoxiang 1 10.82 82 7.56
平均值Mean 11.18 Ab 80 Aa 7.69 Aab
年度间 Between years ns ns ns

表8

不同类型常规粳稻的RVA谱特征值"

品种类型
Type
品种名称
Cultivar
峰值黏度
Peak viscosity
热浆黏度
Trough viscosity
最终黏度
Final viscosity
崩解值
Breakdown
消减值
Setback
回复值
Consistence
2017
高产优质
HG
南粳9108 Nanjing 9108 2109 957 1729 1152 -380 772
南粳505 Nanjing 505 2143 1066 1673 1077 -470 607
徐稻9号Xudao 9 2244 1251 2037 993 -207 786
平均值Mean 2165 Bb 1091 Bc 1813 Bb 1074 Aa -352 Bb 722 Bc
高产不优质
HB
武运5051 Wuyun 5051 2315 1408 2452 907 137 1044
扬粳1612 Yangjing 1612 2350 1464 2475 886 125 1011
华粳8号Huajing 8 2245 1405 2455 840 210 1050
平均值Mean 2303 ABb 1426 ABb 2461 Aa 878 Ab 157 Aa 1035 Aa
中产优质
MG
福粳1606 Fujing 1606 2872 1695 2665 1177 -207 970
苏香粳3号Suxiangjing 3 2600 1656 2495 944 -105 839
松早香1号 Songzaoxiang 1 2623 1628 2445 995 -178 817
平均值Mean 2698 Aa 1660 Aa 2535 Aa 1039 Aa -163 Bb 875 ABb
品种类型
Type
品种名称
Cultivar
峰值黏度
Peak viscosity
热浆黏度
Trough viscosity
最终黏度
Final viscosity
崩解值
Breakdown
消减值
Setback
回复值
Consistence
2018
高产优质
HG
南粳9108 Nanjing 9108 1910 797 1618 1113 -292 821
南粳505 Nanjing 505 2449 1319 1774 1130 -575 555
徐稻9号Xudao 9 2413 1536 2102 877 -311 568
平均值Mean 2257 Aa 1217 Aa 1831 Ab 1040 Aa -393 Bb 648 Bb
高产不优质
HB
武运5051 Wuyun 5051 2451 1649 2658 802 207 1009
扬粳1612 Yangjing 1612 2565 1625 2633 940 68 1008
华粳8号Huajing 8 2178 1381 2378 797 200 997
平均值Mean 2398 Aa 1552 Aa 2556 Aa 846 Ab 158 Aa 1005 Aa
中产优质
MG
福粳1606 Fujing 1606 2718 1551 2338 1167 -380 787
苏香粳3号Suxiangjing 3 2484 1349 2177 1135 -307 828
松早香1号 Songzaoxiang 1 2764 1848 2558 916 -206 710
平均值Mean 2655 Aa 1583 Aa 2358 Aa 1073 Aa -298 Bb 775 ABb
年度间 Between years ns ns ns ns ns ns

表9

2017年不同类型常规粳稻生育期及灌浆结实期温光"

品种类型
Type
品种名称
Cultivar
灌浆结实期
Grain filling period
灌浆结实期温光
Temperature and light in grain filling period
抽穗期Jointing
(month/day)
成熟期Heading
(month/day)
天数
Days
(d)
日平均温度
Mean daily temperature
(℃)
积温
Accumulated temperature
(℃)
日照时数Sunshine hours (h) 降雨量
Rainfall
(mm)
高产优质
HG
南粳9108 Nanjing 9108 8/25 10/21 57 20.77 624.90 215.60 444.50
南粳505 Nanjing 505 8/20 10/25 66 21.06 720.70 256.30 464.20
徐稻9号Xudao 9 8/28 10/25 58 20.38 644.90 214.60 444.50
平均值Mean 60 Aab 20.74 Aa 663.50 Aab 228.83 Aab 451.07 Aa
高产不优质
HB
武运5051 Wuyun 5051 8/22 10/24 63 20.93 699.70 236.10 464.20
扬粳1612 Yangjing 1612 8/25 10/28 63 20.25 676.40 246.80 444.50
华粳8号Huajing 8 8/22 10/25 64 20.85 705.10 239.50 464.20
平均值Mean 63 Aa 20.68 Aa 693.73 Aa 240.80 Aa 457.63 Aa
中产优质
MG
福粳1606 Fujing 1606 8/26 10/19 54 20.45 603.61 201.30 425.00
苏香粳3号Suxiangjing 3 8/18 10/11 54 20.29 598.85 214.80 418.60
松早香1号 Songzaoxiang 1 8/20 10/9 50 20.15 517.65 205.70 396.10
平均值Mean 53 Ab 20.30 Aa 573.37 Ab 207.27 Ab 413.23 Ab
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