[1] FAO. World Food and Agriculture - Statistical Yearbook 2024. Rome: Food and Agriculture Organization of the United Nations, 2024.

[2] 罗锡文, 王在满, 曾山, 等. 水稻机械化直播技术研究进展. 华南农业大学学报, 2019, 40(5): 1–13.

Luo X W, Wang Z M, Zeng S, et al. Recent advances in mechanized direct seeding technology for rice. J South China Agric Univ, 2019, 40(5): 1–13 (in Chinese with English abstract).

[3] Negi P, Rane J, Wagh R S, et al. Direct-seeded rice: genetic improvement of game-changing traits for better adaption. Rice Sci, 2024, 31: 417–433.

[4] Zhang W M, Cheng X Z, Fang D, et al. AT-hook motif nuclear localized (AHL) proteins of ancient origin radiate new functions. Int J Biol Macromol, 2022, 214: 290–300.

[5] Fujimoto S, Matsunaga S, Yonemura M, et al. Identification of a novel plant MAR DNA binding protein localized on chromosomal surfaces. Plant Mol Biol, 2004, 56: 225–239.

[6] Derelle E, Ferraz C, Rombauts S, et al. Genome analysis of the smallest free-living eukaryote Ostreococcus tauri unveils many unique features. Proc Natl Acad Sci USA, 2006, 103: 11647–11652.

[7] Aravind L, Landsman D. AT-hook motifs identified in a wide variety of DNA-binding proteins. Nucleic Acids Res, 1998, 26: 4413–4421.

[8] Širl M, Šnajdrová T, Gutiérrez-Alanís D, et al. AT-hook motif nuclear localised protein 18 as a novel modulator of root system architecture. Int J Mol Sci, 2020, 21: 1886.

[9] Zeng Q K, Song L, Xia M Z, et al. Overexpression of AHL proteins enhances root hair production by altering the transcription of RHD6-downstream genes. Plant Direct, 2023, 7: e517.

[10] Seo M, Lee J Y. Dissection of functional modules of AT-hook motif nuclear localized protein 4 in the development of the root xylem. Front Plant Sci, 2021, 12: 632078.

[11] Cai G Q, Kim S C, Li J W, et al. Transcriptional regulation of lipid catabolism during seedling establishment. Mol Plant, 2020, 13: 984–1000.

[12] Lee K, Seo P J. Coordination of matrix attachment and ATP-dependent chromatin remodeling regulate auxin biosynthesis and Arabidopsis hypocotyl elongation. PLoS One, 2017, 12: e0181804.

[13] Xiao C W, Chen F L, Yu X H, et al. Over-expression of an AT-hook gene, AHL22 delays flowering and inhibits the elongation of the hypocotyl in Arabidopsis thaliana. Plant Mol Biol, 2009, 71: 39–50.

[14] Wong M M, Bhaskara G B, Wen T N, et al. Phosphoproteomics of Arabidopsis Highly ABA-Induced1 identifies AT-Hook-Like10 phosphorylation required for stress growth regulation. Proc Natl Acad Sci USA, 2019, 116: 2354–2363.

[15] Zhou J, Wang X, Lee J Y, et al. Cell-to-cell movement of two interacting AT-hook factors in Arabidopsis root vascular tissue patterning. Plant Cell, 2013, 25: 187–201.

[16] Karami O, Rahimi A, Khan M, et al. A suppressor of axillary meristem maturation promotes longevity in flowering plants. Nat Plants, 2020, 6: 368–376.

[17] Rahimi A, Karami O, Balazadeh S, et al. miR156-independent repression of the ageing pathway by longevity-promoting AHL proteins in Arabidopsis. New Phytol, 2022, 235: 2424–2438.

[18] Rahimi A, Karami O, Lestari A D, et al. Control of cambium initiation and activity in Arabidopsis by the transcriptional regulator AHL15. Curr Biol, 2022, 32: 1764–1775.e3.

[19] Liu H, Hu D X, Du P X, et al. Single-cell RNA-seq describes the transcriptome landscape and identifies critical transcription factors in the leaf blade of the allotetraploid peanut (Arachis hypogaea L.). Plant Biotechnol J, 2021, 19: 2261–2276.

[20] Rayapuram N, Jarad M, Alhoraibi H M, et al. Chromatin phosphoproteomics unravels a function for AT-hook motif nuclear localized protein AHL13 in PAMP-triggered immunity. Proc Natl Acad Sci USA, 2021, 118: e2004670118.

[21] Jin Y, Luo Q, Tong H N, et al. An AT-hook gene is required for Palea formation and floral organ number control in rice. Dev Biol, 2011, 359: 277–288.

[22] Uzair M, Xu D W, Schreiber L, et al. PERSISTENT TAPETAL CELL2 is required for normal tapetal programmed cell death and pollen wall patterning. Plant Physiol, 2020, 182: 962–976.

[23] Zhou L G, Liu Z C, Liu Y H, et al. A novel gene OsAHL1 improves both drought avoidance and drought tolerance in rice. Sci Rep, 2016, 6: 30264.

[24] Liu X Y, Gao Y X, Guo Z Q, et al. MoIug4 is a novel secreted effector promoting rice blast by counteracting host OsAHL1-regulated ethylene gene transcription. New Phytol, 2022, 235: 1163–1178.

[25] Chen C J, Wu Y, Li J W, et al. TBtools-II: a “one for all, all for one” bioinformatics platform for biological big-data mining. Mol Plant, 2023, 16: 1733–1742.

[26] 曾栋昌, 马兴亮, 谢先荣, 等. 植物CRISPR/Cas9多基因编辑载体构建和突变分析的操作方法. 中国科学(生命科学), 2018, 48: 783–794.

Zeng D C, Ma X L, Xie X R, et al. A protocol for CRISPR/Cas9-based multi-gene editing and sequence decoding of mutant sites in plants. Sci Sin Vitae, 2018, 48: 783–794 (in Chinese with English abstract).

[27] Liu Q, Wang C, Jiao X Z, et al. Hi-TOM: a platform for high-throughput tracking of mutations induced by CRISPR/Cas systems. Sci China Life Sci, 2019, 62: 1–7.

[28] Wang S, Liu W N, He Y, et al. bZIP72 promotes submerged rice seed germination and coleoptile elongation by activating ADH1. Plant Physiol Biochem, 2021, 169: 112–118.

[29] Fu J, Wu H, Ma S Q, et al. OsJAZ1 attenuates drought resistance by regulating JA and ABA signaling in rice. Front Plant Sci, 2017, 8: 2108.

[30] Lin C C, Chao Y T, Chen W C, et al. Regulatory cascade involving transcriptional and N-end rule pathways in rice under submergence. Proc Natl Acad Sci USA, 2019, 116: 3300–3309.

[31] Reed R C, Bradford K J, Khanday I. Seed germination and vigor: ensuring crop sustainability in a changing climate. Heredity, 2022, 128: 450–459.

[32] Wasternack C, Song S S. Jasmonates: biosynthesis, metabolism, and signaling by proteins activating and repressing transcription. J Exp Bot, 2017, 68: 1303–1321.

[33] Ruan J J, Zhou Y X, Zhou M L, et al. Jasmonic acid signaling pathway in plants. Int J Mol Sci, 2019, 20: 2479.

[34] Dave A, Hernández M L, He Z S, et al. 12-oxo-phytodienoic acid accumulation during seed development represses seed germination in Arabidopsis. Plant Cell, 2011, 23: 583–599.

[35] Linkies A, Leubner-Metzger G. Beyond gibberellins and abscisic acid: how ethylene and jasmonates control seed germination. Plant Cell Rep, 2012, 31: 253–270.