[1] Panni M K, Khan N U, Fitmawati, et al. Heterotic studies and inbreeding depression in F₂ populations of upland cotton. Pak J Bot, 2012, 44: 1013–1020.

[2] Maryum Z, Luqman T, Nadeem S, et al. An overview of salinity stress, mechanism of salinity tolerance and strategies for its management in cotton. Front Plant Sci, 2022, 13: 907937.

[3] 毛树春, 马小艳, 程思贤, 等. 我国高品质棉花产需分析与发展建议. 中国棉花, 2020, 47(3): 1–5.
Mao S C, Ma X Y, Cheng S X, et al. Analysis and strategy for the production and demand of high quality cotton in China. China Cotton, 2020, 47(3): 1–5 (in Chinese with English abstract).

[4] 阿里甫·艾尔西, 朱家辉, 李进, 等. 新疆长绒棉育种现状、趋势及研究进展. 新疆农业科学, 2020, 57: 393–400.
Alifu Aierxi, Zhu J H, Li J, et al. Progress, problems and prospects of Xinjiang long-staple cotton (G. barbadense L.) breeding. Xinjiang Agric Sci, 2020, 57: 393–400 (in Chinese with English abstract).

[5] 洪梅, 张磊磊, 陈晋瑞, 等. 新疆长绒棉机采的实践与思考. 中国棉花, 2020, 47(11): 8–10.
Hong M, Zhang L L, Chen J R, et al. Practice and thinking on mechanical harvest of long staple cotton in Xinjiang. China Cotton, 2020, 47(11): 8–10 (in Chinese).

[6] Wang Y X, Lyu Q Q, Tian J S, et al. The high genetic yield of Xinjiang cotton is associated with improvements in boll-leaf system photosynthesis. Field Crops Res, 2023, 304: 109176.

[7] 郑子漂, 徐海江, 林涛, 等. 新疆长绒棉育成品种演变趋势及综合评价. 中国农业大学学报, 2022, 27(6): 55–70.

Zheng Z P, Xu H J, Lin T, et al. Evolution tendency and comprehensive evaluation of long-staple cotton bred varieties in Xinjiang. J Chin Agric Univ, 2022, 27(6): 55–70 (in Chinese with English abstract).

[8] Saha S, Wu J, Jenkins J N, et al. Interspecific chromosomal effects on agronomic traits in Gossypium hirsutum by AD analysis using intermated G. barbadense chromosome substitution lines. Theor Appl Genet, 2013, 126: 109–117.

[9] Witt T W, Ulloa M, Schwartz R C, et al. Response to deficit irrigation of morphological, yield and fiber quality traits of upland (Gossypium hirsutum L.) and Pima (G. barbadense L.) cotton in the Texas High Plains. Field Crops Res, 2020, 249: 107759.

[10] Holladay S K, Bridges W, Jones M A, et al. Growth, boll development, agronomic performance, and fiber quality of Gossypium barbadense L. in the southeastern U.S. Coastal Plain. Crop Sci, 2022, 62: 1461–1471.

[11] 新疆维吾尔自治区统计局. 新疆统计年鉴. 北京: 中国统计出版社, 2024.
Statistics Bureau of Xinjiang Uygur Autonomous Region. Xinjiang Statistical Yearbook. Beijing: China Statistics Press, 2024 (in Chinese). 

[12] 孔杰, 王为然, 朱家辉, 等. 新疆海岛棉主要育成品种性状特征与系谱来源分析. 新疆农业科学, 2018, 55: 589–600.
Kong J, Wang W R, Zhu J H, et al. Characteristics and pedigree analysis of main cultivated varieties of Xinjiang sea-island cotton (G. barbadense L.). Xinjiang Agric Sci, 2018, 55: 589–600 (in Chinese with English abstract).

[13] 王丽, 张巨松, 宁丽云, 等. 施氮量和种植密度互作对海岛棉冠层光分布、产量及氮素利用的影响. 植物营养与肥料学报, 2025, 31: 1998–2014.

Wang L, Zhang J S, Ning L Y, et al. Interactive effects of planting density and nitrogen application rate on canopy light distribution, yield and nitrogen use of sea island cotton. J Plant Nutr Fert, 2025, 31: 1998–2014 (in Chinese with English abstract).

[14] Gifford R M, Thorne J H, Hitz W D, et al. Crop productivity and photoassimilate partitioning. Science, 1984, 225: 801–808.

[15] Murchie E H, Reynolds M, Slafer G A, et al. A ‘wiring diagram’ for source strength traits impacting wheat yield potential. J Exp Bot, 2023, 74: 72–90.

[16] Elmore C D. Contributions of the capsule wall and bracts to the developing cotton Fruit. Crop Sci, 1973, 13: 751–752.

[17] Yan P, Li X J, Shan W H, et al. Plasticity of sympodial leaves in the cotton boll-leaf system governs single boll weight by modulating photosynthetic assimilate supply. Plant J, 2026, 125: e70729.

[18] Zhan D X, Yang Y, Hu Y Y, et al. Contributions of nonleaf organs to the yield of cotton grown with different water supply. Sci World J, 2014, 2014: 602747.

[19] Liang F B, Chen M Z, Shi Y, et al. Single boll weight depends on photosynthetic function of boll-leaf system in field-grown cotton plants under water stress. Photosynth Res, 2021, 150: 227–237.

[20] Wullschleger S D, Oosterhuis D M. Photosynthetic carbon production and use by developing cotton leaves and bolls. Crop Sci, 1990, 30: 1259–1264.

[21] Wang Y X, Chen M Z, Liang F B, et al. Photosynthates competition within the boll-leaf system is alleviated with the improvement of photosynthetic performance during the succession of Xinjiang cotton cultivars. Ind Crops Prod, 2021, 160: 113121.

[22] Hu Y Y, Zhang Y L, Luo H H, et al. Important photosynthetic contribution from the non-foliar green organs in cotton at the late growth stage. Planta, 2012, 235: 325–336.

[23] Zou J, Hu W, Loka D A, et al. Carbon assimilation and distribution in cotton photosynthetic organs is a limiting factor affecting boll weight formation under drought. Front Plant Sci, 2022, 13: 1001940.

[24] Tang F Y, Xiao W J. Dry matter accumulation and partitioning in various fractions of cotton bolls. Exp Agric, 2013, 49: 543–555.

[25] Zhang Q P, Luo D, Sun Y Y, et al. Cotton harvest aids promote the translocation of bur-stored photoassimilates to enhance single boll weight. Ind Crops Prod, 2023, 195: 116375.

[26] Chen W, Yao J B, Li Y, et al. Nulliplex-branch, a TERMINAL FLOWER 1 ortholog, controls plant growth habit in cotton. Theor Appl Genet, 2019, 132: 97–112.

[27] Chao L, Pan Z Y, Wang J, et al. Genetic mapping and analysis of a compact plant architecture and precocious mutant in upland cotton. Plants, 2022, 11: 1483.

[28] Lamlom S F, Yehia W M B, Kotb H M K, et al. Genetic improvement of Egyptian cotton (Gossypium barbadense L.) for high yield and fiber quality properties under semi arid conditions. Sci Rep, 2024, 14: 7723.

[29] Meng Q Y, Xie P H, Xu Z P, et al. Pangenome analysis reveals yield- and fiber-related diversity and interspecific gene flow in Gossypium barbadense L. Nat Commun, 2025, 16: 4995.

[30] Han J M, Flexas J, Xiong D L, et al. Regulation of photosynthesis and water-use efficiency in Pima and upland cotton species subjected to drought and recovery. Photosynthetica, 2024, 62: 6–15.

[31] 姚贺盛, 张亚黎, 易小平, 等. 海岛棉和陆地棉叶片光合特性、冠层结构及物质生产的差异. 中国农业科学, 2015, 48: 251–261.
Yao H S, Zhang Y L, Yi X P, et al. Study on differences in comparative canopy structure characteristics and photosynthetic carbon assimilation of field-grown Pima cotton (Gossypium barbadense) and upland cotton (G. hirsutum). Sci Agric Sin, 2015, 48: 251–261 (in Chinese with English abstract).

[32] 张亚黎, 姚贺盛, 罗毅, 等. 海岛棉和陆地棉叶片光合能力的差异及限制因素. 生态学报, 2011, 31: 1803–1810.
Zhang Y L, Yao H S, Luo Y, et al. Difference in leaf photosynthetic capacity between Pima cotton (Gossypium barbadense) and upland cotton (G.hirsutum) and analysis of potential constraints. Acta Ecol Sin, 2011, 31: 1803–1810 (in Chinese with English abstract).

[33] Han J M, Zhang Y J, Lei Z Y, et al. The higher area-based photosynthesis in Gossypium hirsutum L. is mostly attributed to higher leaf thickness. Photosynthetica, 2019, 57: 420–427.

[34] Lu Z M, Chen J W, Percy R G, et al. Photosynthetic rate, stomatal conductance and leaf area in two cotton species (Gossypium barbadense and Gossypium hirsutum) and their relation with heat resistance and yield. Aust J Plant Physiol, 1997, 24: 693–700.

[35] Wise R. A comparison of leaf anatomy in field-grown Gossypium hirsutum and G. barbadense. Ann Bot, 2000, 86: 731–738.

[36] Zhang D M, Luo Z, Liu S H, et al. Effects of deficit irrigation and plant density on the growth, yield and fiber quality of irrigated cotton. Field Crops Res, 2016, 197: 1–9.

[37] Sinclair T R, Rufty T W, Lewis R S. Increasing photosynthesis: unlikely solution for world food problem. Trends Plant Sci, 2019, 24: 1032–1039.

[38] Chen M Z, Zhang Y L, Liang F B, et al. The net photosynthetic rate of the cotton boll-leaf system determines boll weight under various plant densities. Eur J Agron, 2021, 125: 126251.

[39] Chen M Z, Liang F B, Yan Y H, et al. Boll-leaf system gas exchange and its application in the analysis of cotton photosynthetic function. Photosynth Res, 2021, 150: 251–262.

[40] Araus J L, Sanchez-Bragado R, Vicente R. Improving crop yield and resilience through optimization of photosynthesis: Panacea or pipe dream? J Exp Bot, 2021, 72: 3936–3955.

[41] Smith E N, van Aalst M, Tosens T, et al. Improving photosynthetic efficiency toward food security: Strategies, advances, and perspectives. Mol Plant, 2023, 16: 1547–1563.

[42] Croce R, Carmo-Silva E, Cho Y B, et al. Perspectives on improving photosynthesis to increase crop yield. Plant Cell, 2024, 36: 3944–3973.

[43] Shi W J, Xiao G, Struik P C, et al. Quantifying source-sink relationships of rice under high night-time temperature combined with two nitrogen levels. Field Crops Res, 2017, 202: 36–46.

[44] Xing F F, Han Y C, Feng L, et al. Genotypic variation in spatiotemporal distribution of canopy light interception in relation to yield formation in cotton. J Cotton Res, 2018, 1: 13.

[45] Nie J J, Qin D L, Mao L L, et al. Genotypic variance in ¹³C-photosynthate partitioning and within-plant boll distribution in cotton. J Cotton Res, 2020, 3: 15.

[46] Yan Y Y, Duan F Y, Li X, et al. Photosynthetic capacity and assimilate transport of the lower canopy influence maize yield under high planting density. Plant Physiol, 2024, 195: 2652–2667.

[47] Yang L Y, Duan J H, Liu Y, et al. Changes in carbohydrate distribution in cotton photosynthetic organs and increase in boll weight reduce yield loss under high temperature. J Exp Bot, 2024, 75: 3483–3499.

[48] 曹新川, 胡守林, 韩秀锋,等. 海岛棉棉铃阶段性发育与产量品质的关系. 作物学报, 2020, 46: 300–306.

Cao X C, Hu S L, Han X F, et al. Relationship of stage development of cotton bolls with yield and quality in island cotton. Acta Agron Sin, 2020, 46: 300–306 (in Chinese with English abstract).

[49] Tang F Y, Wang T, Zhu J M. Carbohydrate profiles during cotton (Gossypium hirsutum L.) boll development and their relationships to boll characters. Field Crops Res, 2014, 164: 98–106.

[50] Le Y, Chen M L, Zhu D, et al. Transcriptional Regulation of SWEET15_A01 by MYB44/bHLH3 Modulates Carbon Allocation in Cotton Ovule and Fibre to Affect Seed and Fibre Traits. Plant Biotechnol J, 2026, 24: 1769–1791.

[51] Wang Y H, Feng Y, Xu N Y, et al. Response of the enzymes to nitrogen applications in cotton fiber (Gossypium hirsutum L.) and their relationships with fiber strength. Sci China Ser C-Life Sci, 2009, 52: 1065–1072

[52] Li, H J, Wang, J W, Huang X L, et al. Phosphate fertilization promotes fiber elongation by affecting sucrose inversion, K^＋accumulation, and malate synthesis in cotton fiber. Field Crops Res, 2023, 303, 109119.

[53] Jin K N, Chen G X, Yang Y R, et al. Strategies for manipulating Rubisco and creating photorespiratory bypass to boost C₃ photosynthesis: Prospects on modern crop improvement. Plant Cell Environ, 2023, 46: 363–378.

[54] Yang Y P, Wu K F, Yu D Q, et al. Inducer of CBF Expression1 (ICE1) interacts with WRKY46 to modulate salicylic acid-induced leaf senescence in Arabidopsis. Plant Cell Environ, 2025, 48: 6140–6152.

[55] 刘韬奋, 罗单, 张启鹏, 等. 乙烯利催熟对机采棉铃重和纤维品质的影响. 作物学报, 2024, 50: 209–218.

Liu T F, Luo D, Zhang Q P, et al. Ethephon ripening affects boll weight and fiber quality of machine-harvested cotton. Acta Agron Sin, 2024, 50: 209–218 (in Chinese with English abstract).

[56] Brookbank B P, Patel J, Gazzarrini S, et al. Role of basal ABA in plant growth and development. Genes, 2021, 12: 1936.

[57] Guo Y T, Zou L M, Huang L H, et al. Overexpression of GhGGPPS10 and GhGGPPS23 enhances plant biomass accumulation and optimizes photosynthetic performance in cotton. Plant Cell Environ, 2026, 49: 700–713.

[58] Zhou J Y, Nie J J, Kong X Q, et al. Cotton yield stability achieved through manipulation of vegetative branching and photoassimilate partitioning under reduced seedling density and double seedlings per hole. Field Crops Res, 2023, 303: 109117.

[59] Wu F Q, Tang Q X, Cui J P, et al. Deficit irrigation combined with a high planting density optimizes root and soil water–nitrogen distribution to enhance cotton productivity in arid regions. Field Crops Res, 2024, 317: 109524.

[60] Zuo W Q, Wu B J, Wang Y X, et al. Optimal row spacing configuration to improve cotton yield or quality is regulated by plant density and irrigation rate. Field Crops Res, 2024, 305: 109187.

[61] 王士红, 杨中旭, 史加亮, 等. 增密减氮对棉花干物质和氮素积累分配及产量的影响. 作物学报, 2020, 46: 395–407.

Wang S H, Yang Z X, Shi J L, et al. Effects of increasing planting density and decreasing nitrogen rate on dry matter, nitrogen accumulation and distribution, and yield of cotton. Acta Agron Sin, 2020, 46: 395–407 (in Chinese with English abstract).

[62] Ma Y, Ye Z P, Li W, et al. Micronutrient uptake, partitioning and accumulation in cotton under drip irrigation with mulch in response to phosphorus application in root morphology changes. Field Crops Res, 2026, 336: 110234.

[63] Wang C, Wang S S, Zhang Y T, et al. Optimizing photoassimilate synthesis, metabolism, and loading in cotton’s “source-sink-flow” system by boron fertilizer management to increase boll weight. Crop J, 2026, 14: 1028–1038.