[1] Wang X, Liu Y, Ouyang L, Yao R N, He D L, Han Z K, Li W T, Ding Y B, Wang Z H, Kang Y P, Yan L Y, Chen Y N, Huai D X, Jiang H F, Lei Y, Liao B S. Metabolomics combined with transcriptomics analyses of mechanism regulating testa pigmentation in peanut. Front Plant Sci, 2022, 13: 1065049.

[2] 李佳伟, 马钰聪, 杨鑫雷, 王梅, 崔顺立, 侯名语, 刘立峰, 胡梦蝶, 蒋晓霞, 穆国俊. 花生种皮色素合成相关通路的转录组-代谢组学联合分析. 植物遗传资源学报, 2022, 23: 240–254.

Li J W, Ma Y C, Yang X L, Wang M, Cui S L, Hou M Y, Liu L F, Hu M D, Jiang X X, Mu G J. Transcriptomics-metabolomics combined analysis highlight the mechanism of testa pigment formation in peanut (Arachis hypogaea L.). J Plant Genet Resour, 2022, 23: 240–254 (in Chinese with English abstract).

[3] Zhang K, Yuan M, Xia H, He L Q, Ma J, Wang M X, Zhao H L, Hou L, Zhao S Z, Li P C, Tian R Z, Pan J W, Li G H, Thudi M, Ma C L, Wang X J, Zhao C Z. BSA-seq and genetic mapping reveals AhRt2 as a candidate gene responsible for red testa of peanut. Theor Appl Genet, 2022, 135: 1529–1540.

[4] Chen L, Yan F F, Chen W B, Zhao L, Zhang J L, Lu Q, Liu R. Procyanidin from peanut skin induces antiproliferative effect in human prostate carcinoma cells DU145. Chem Biol Interact, 2018, 288: 12–23.

[5] Zhu F. Anthocyanins in cereals: Composition and health effects. Food Res Int, 2018, 109: 232–249.

[6] Tsuda T. Dietary anthocyanin-rich plants: Biochemical basis and recent progress in health benefits studies. Mol Nutr Food Res, 2012, 56: 159–170.

[7] Attree R, Du B, Xu B J. Distribution of phenolic compounds in seed coat and Cotyledon, and their contribution to antioxidant capacities of red and black seed coat peanuts (Arachis hypogaea L.). Ind Crops Prod, 2015, 67: 448–456.

[8] Alseekh S, Perez de Souza L, Benina M, Fernie A R. The style and substance of plant flavonoid decoration; towards defining both structure and function. Phytochemistry, 2020, 174: 112347.

[9] Iwashina T. Flavonoid function and activity to plants and other organisms. Biol Sci Space, 2003, 17: 24–44.

[10] Tanaka Y, Brugliera F, Chandler S. Recent progress of flower colour modification by biotechnology. Int J Mol Sci, 2009, 10: 5350–5369. 

[11] Wen W W, Alseekh S, Fernie A R. Conservation and diversification of flavonoid metabolism in the plant kingdom. Curr Opin Plant Biol, 2020, 55: 100–108.

[12] Wang X, Zhang X C, Hou H X, Ma X, Sun S L, Wang H W, Kong L R. Metabolomics and gene expression analysis reveal the accumulation patterns of phenylpropanoids and flavonoids in different colored-grain wheats (Triticum aestivum L.). Food Res Int, 2020, 138: 109711.

[13] Grotewold E. The genetics and biochemistry of floral pigments. Annu Rev Plant Biol, 2006, 57: 761–780.

[14] Duan H R, Wang L R, Cui G X, Zhou X H, Duan X R, Yang H S. Identification of the regulatory networks and hub genes controlling alfalfa floral pigmentation variation using RNA-sequencing analysis. BMC Plant Biol, 2020, 20: 110.

[15] 吴紫萱, 薛其勤, 杨会, 刘风珍. 花生种皮颜色研究进展. 山东农业科学, 2022, 54(1): 152–156.

Wu Z X, Xue Q Q, Yang H, Liu F Z. Research progress on testa color of peanut. Shandong Agric Sci, 2022, 54(1): 152–156 (in Chinese with English abstract).

[16] Hu M D, Li J W, Hou M Y, Liu X Q, Cui S L, Yang X L, Liu L F, Jiang X X, Mu G J. Transcriptomic and metabolomic joint analysis reveals distinct flavonoid biosynthesis regulation for variegated testa color development in peanut (Arachis hypogaea L.). Sci Rep, 2021, 11: 10721.

[17] Huang J Y, Xing M H, Li Y, Cheng F, Gu H H, Yue C P, Zhang Y J. Comparative transcriptome analysis of the skin-specific accumulation of anthocyanins in black peanut (Arachis hypogaea L.). J Agric Food Chem, 2019, 67: 1312−1324.

[18] Wan L Y, Li B, Lei Y, Yan L Y, Huai D X, Kang Y P, Jiang H F, Tan J Z, Liao B S. Transcriptomic profiling reveals pigment regulation during peanut testa development. Plant Physiol Biochem, 2018, 125: 116–125.

[19] Lou Q, Liu Y L, Qi Y Y, Jiao S Z, Tian F F, Jiang L, Wang Y J. Transcriptome sequencing and metabolite analysis reveals the role of delphinidin metabolism in flower colour in grape hyacinth. J Exp Bot, 2014, 65: 3157–3164.

[20] Wu Q, Wu J, Li S S, Zhang H J, Feng C Y, Yin D D, Wu R Y, Wang L S. Transcriptome sequencing and metabolite analysis for revealing the blue flower formation in waterlily. BMC Genomics, 2016, 17: 897.

[21] Xia H, Zhu L, Zhao C Z, Li K, Shang C L, Hou L, Wang M X, Shi J, Fan S J, Wang X J. Comparative transcriptome analysis of anthocyanin synthesis in black and pink peanut. Plant Signal Behav, 2020, 15: 1721044.

[22] Koes R, Verweij W, Quattrocchio F. Flavonoids: a colorful model for the regulation and evolution of biochemical pathways. Trends Plant Sci, 2005, 10: 236–242.

[23] Labbé D, Provencal M, Lamy S, Boivin D, Gingras D, Béliveau R. The flavonols quercetin, kaempferol, and myricetin inhibit hepatocyte growth factor-induced medulloblastoma cell migration. J Nutr, 2009, 139: 646–652.

[24] Kuang Q J, Yu Y Y, Attree R, Xu B J. A comparative study on anthocyanin, saponin, and oil profiles of black and red seed coat peanut (Arachis hypogacea) grown in China. Int J Food Prop, 2017, 20: S131–S140.

[25] Nabavi S M, Šamec D, Tomczyk M, Milella L, Russo D, Habtemariam S, Suntar I, Rastrelli L, Daglia M, Xiao J B, Giampieri F, Battino M, Sobarzo-Sanchez E, Nabavi S F, Yousefi B, Jeandet P, Xu S W, Shirooie S. Flavonoid biosynthetic pathways in plants: Versatile targets for metabolic engineering. Biotechnol Adv, 2020, 38: 107316.

[26] Zhou C B, Mei X, Rothenberg D O, Yang Z B, Zhang W T, Wan S H, Yang H J, Zhang L Y. Metabolome and transcriptome analysis reveals putative genes involved in anthocyanin accumulation and coloration in white and pink tea (Camellia sinensis) flower. Molecules, 2020, 25: 190.

[27] Khlestkina E K, Shoeva O Y, Gordeeva E I. Flavonoid biosynthesis genes in wheat. Russ J Genet Appl Res, 2015, 5: 268–278.

[28] 苏俏, 金欣欣, 李玉荣, 程增书, 宋亚辉, 杨永庆, 王瑾. 影响多彩花生种皮颜色的关键代谢物及ANS基因分析. 华北农学报, 2022, 37(增刊): 19–25.

Su Q, Jin X X, Li Y R, Cheng Z S, Song Y H, Yang Y Q, Wang J. Analysis of key metabolites and ANS genes affecting seed testa color of peanut. Acta Agric Boreali-Sin, 2022, 37(S1): 19–25 (in Chinese with English abstract).

[29] Zhao Z L, Wu M, Zhan Y L, Zhan K H, Chang X L, Yang H S, Li Z M. Characterization and purification of anthocyanins from black peanut (Arachis hypogaea L.) skin by combined column chromatography. J Chromatogr A, 2017, 1519: 74–82.