[1]Ishiguro S, Nakamura K. Characterization of a cDNA encoding a novel DNA-binding protein, SPF1, that recognizes SP8 sequences in the 5' upstream regions of genes coding for sporamin and β-amylase from sweet potato. Mol Gen Genet, 1994, 244: 563–571



[2]Eulgem T, Rushton PJ, Robatzek S, Somssich I E. The WRKY super family of plant transcription factors. Trends Plant Sci, 2000, 5: 199–206



[3]Yu L, Chen C, Chen Z. Evidence for an important role of the WRKY DNA-binding proteins in the regulation of the NPR1 gene expression. Plant Cell, 2001, 13: 1527–1539



[4]Dong J, Chen C, Chen Z. Expression profiles of the Arabidopsis WRKY gene superfamily during plant defense response. Plant Mol Biol, 2003, 51: 21–37



[5]Jiang Y, Deyholos M K. Comprehensive transcriptional profiling of NaCl-stressed Arabidopsis roots reveals novel classes of responsive genes. BMC Plant Biol, 2006, 6: 25



[6]Jiang Y, Deyholos M K. Functional characterization of Arabidopsis NaCl-inducible WRKY25 and WRKY33 transcription factors in abiotic stresses. Plant Mol Biol, 2009, 69: 91–105



[7]Zou C, Jiang W, Yu D. Male gametophyte-specific WRKY34 transcription factor mediates cold sensitivity of mature pollen in Arabidopsis. J Exp Bot, 2010, 61: 3901–3914



[8]Mao G, Meng X, Liu Y, Zheng Z, Chen Z, Zhang S. Phosphorylation of a WRKY transcription factor by two pathogen-responsive MAPKs drives phytoalexin biosynthesis in Arabidopsis. Plant Cell, 2011, 23: 1639–1653



[9]Luo M, Dennis ES, Berger F, Peacock WJ, Chaudhury A. MINISEED3 (MINI3), a WRKY family gene, and HAIKU2 (IKU2), a leucine-rich repeat (LRR) KINASE gene, are regulators of seed size in Arabidopsis. Proc Natl Acad Sci USA, 2005, 102: 17531–17536



[10]Xie Z, Zhang Z L, Zou X, Huang J, Ruas P, Thompson D, Shen Q J. Annotations and functional analysis of the rice WRKY gene superfamily reveal positive and negative regulators of abscisic acid signaling in aleurone cells. Plant Physiol, 2005, 137: 176–189



[11]Ross C A, Liu Y, Shen Q J. The WRKY gene family in rice (Oryza sativa). J Integr Plant Biol, 2007, 49: 827–842



[12]Ramamoorthy R, Jiang S Y, Kumar N, Venkatesh PN, Ramachandran S. A comprehensive transcriptional profiling of the WRKY gene family in rice under various abiotic and phytohormone treatments. Plant Cell Physiol, 2008, 49: 865–879



[13]Tao Z, Kou Y, Liu H, Li X, Xiao J, Wang S. OsWRKY45 alleles play different roles in abscisic acid signalling and salt stress tolerance but similar roles in drought and cold tolerance in rice. J Exp Bot, 2011, 62: 4863–4874



[14]Wu X, Shiroto Y, Kishitani S, Ito Y, Toriyama K. Enhanced heat and drought tolerance in transgenic rice seedlings overexpressing OsWRKY11 under the control of HSP101 promoter. Plant Cell Rep, 2009, 28: 21–30



[15]Hwang S H, Yie S W, Hwang D J. Heterologous expression of OsWRKY6 gene in Arabidopsis activates the expression of defense related genes and enhances resistance to pathogens. Plant Sci, 2011, 181: 316–323



[16]Song Y, Chen L, Zhang L, Yu D. Overexpression of OsWRKY72 gene interferes in the abscisic acid signal and auxin transport pathway of Arabidopsis. J Biosci, 2010, 35: 459–471



[17]Yin G, Xu H, Xiao S, Qin Y, Li Y, Yan Y, Hu Y. The large soybean (Glycine max) WRKY TF family expanded by segmental duplication events and subsequent divergent selection among subgroups. BMC Plant Biol, 2013, 13: 148



[18]Dou L, Zhang X, Pang C, Song M, Wei H, Fan S, Yu S. Genome-wide analysis of the WRKY gene family in cotton. Mol Genet Genomics, 2014: 1-19



[19]Pnueli L, Hallak-Herr E, Rozenberg M, Cohen M, Goloubinoff P, Kaplan A, Mittler R. Molecular and biochemical mechanisms associated with dormancy and drought tolerance in the desert legume Retama raetam. Plant J, 2002, 31: 319–330



[20]Wei W, Zhang Y, Han L, Guan Z, Chai T. A novel WRKY transcriptional factor from Thlaspi caerulescens negatively regulates the osmotic stress tolerance of transgenic tobacco. Plant Cell Rep, 2008, 27: 795–903



[21]Marè C, Mazzucotelli E, Crosatti C, Francia E, Stanca Am, Cattivelli L. Hv-WRKY38: a new transcription factor involved in cold- and drought-response in barley. Plant Mol Biol, 2004, 55: 339–416



[22]Wang C, Deng P, Chen L, Wang X, Ma H, Hu W, Yao N, Feng Y, Chai R, Yang G, He G. A wheat WRKY transcription factor TaWRKY10 confers tolerance to multiple abiotic stresses in transgenic tobacco. PLoS One, 2013, 8: e65120



[23]Niu C F, Wei W, Zhou Q Y, Tian A G, Hao Y J, Zhang W K, Ma B, Lin Q, Zhang Z B, Zhang J S, Chen S Y. Wheat WRKY genes TaWRKY2 and TaWRKY19 regulate abiotic stress tolerance in transgenic Arabidopsis plants. Plant Cell Environ, 2012, 35: 1156–1170



[24]Ling H Q, Zhao S, Liu D, Wang J, Sun H, Zhang C, Fan H, Li D, Dong L, Tao Y, Gao C, Wu H, Li Y, Cui Y, Guo X, Zheng S, Wang B, Yu K, Liang Q, Yang W, Lou X, Chen J, Feng M, Jian J, Zhang X, Luo G, Jiang Y, Liu J, Wang Z, Sha Y, Zhang B, Wu H, Tang D, Shen Q, Xue P, Zou S, Wang X, Liu X, Wang F, Yang Y, An X, Dong Z, Zhang K, Zhang X, Luo M C, Dvorak J, Tong Y, Wang J, Yang H, Li Z, Wang D, Zhang A, Wang J. Draft genome of the wheat A-genome progenitor Triticum urartu. Nature, 2013, 496: 87–90



[25]Finn R D, Mistry J, Schuster-Böckler B, Griffiths-Jones S, Volker Hollich1 TL, Moxon S, Marshall M, Khanna A, Durbin R, Eddy S R, Sonnhammer E L L, Bateman A. Pfam: clans, web tools and services. Nucl Acids Res, 2006, 34: D247–D251



[26]Letunic I, Doerks T, Bork P. SMART 6: recent updates and new developments. Nucl Acids Res, 2009, 37: D229–D232



[27]Wei H, Li W, Sun X, Zhu S, Zhu J. Systematic analysis and comparison of nucleotide-binding site disease resistance genes in a diploid cotton Gossypium raimondii. PLoS One, 2013, 8: e68435



[28]Gaut B S, Doebley J F. DNA sequence evidence for the segmental allotetraploid origin of maize. Proc Natl Acad Sci USA, 1997, 94: 6809–6814



[29]Suyama M, Torrents D, Bork P. PAL2NAL: robust conversion of protein sequence alignments into the corresponding codon alignments. Nucl Acids Res, 2006, 34: W609–W612



[30]郭安源, 朱其慧, 陈新, 罗静初. GSDS: 基因结构显示系统. 遗传, 2007, 29: 1023-1026



Guo A Y, Zhu Q H, Chen X, Luo J C. GSDS: a gene structure display server. Hereditas (Beijing), 2007, 29: 1023–1026 (in Chinese with English abstract)



[31]Holub E B. The arms race is ancient history in Arabidopsis, the wildflower. Nat Rev Genet, 2001, 2: 516–527



[32]Peng X, Zhao Y, Cao J, Zhang W, Jiang H, Li X, Ma Q, Zhu S, Cheng B. CCCH-type zinc finger family in maize: genome-wide identification, classification and expression profiling under abscisic acid and drought treatments. PLoS One, 2012, 7: e40120



[33]Wu H, Ni Z, Yao Y, Guo G, Sun Q. Cloning and expression profiles of 15 genes encoding WRKY transcription factor in wheat (Triticum aestivem L.). Prog Nat Sci, 2008, 18: 697–705