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作物学报 ›› 2025, Vol. 51 ›› Issue (7): 1801-1813.doi: 10.3724/SP.J.1006.2025.41086

• 作物遗传育种·种质资源·分子遗传学 • 上一篇    下一篇

全基因组关联分析定位与挖掘小麦氮高效基因

赵超男1(), 王金凤1, 张玉1, 张丽1, 李瑞琦1, 王鹏飞1, 李鸽子1, 张宏军2, 虞波1,*(), 康国章1,*()   

  1. 1河南农业大学 / 国家小麦工程技术研究中心, 河南郑州 450046
    2中国农业科学院作物科学研究所 / 作物分子育种国家工程实验室, 北京 100081
  • 收稿日期:2024-12-03 接受日期:2025-04-27 出版日期:2025-07-12 网络出版日期:2025-05-13
  • 通讯作者: *虞波, E-mail: yubo@henau.edu.cn; 康国章, E-mail: guozhangkang@henau.edu.cn
  • 作者简介:E-mail: zhaocn@stu.henau.edu.cn
  • 基金资助:
    国家自然科学基金项目(32401739);神农种业实验室“一流课题”项目(SN01-2022-01)

Genome-wide association study for the identification and characterization of nitrogen efficiency-related genes in wheat

ZHAO Chao-Nan1(), WANG Jin-Feng1, ZHANG Yu1, ZHANG Li1, LI Rui-Qi1, WANG Peng-Fei1, LI Ge-Zi1, ZHANG Hong-Jun2, YU Bo1,*(), KANG Guo-Zhang1,*()   

  1. 1Henan Agricultural University / National Wheat Engineering Research Center, Zhengzhou 450046, Henan, China
    2Institute of Crop Sciences, Chinese Academy of Agricultural Sciences / National Engineering Research Center of Crop Molecular Breeding, Beijing 100081, China
  • Received:2024-12-03 Accepted:2025-04-27 Published:2025-07-12 Published online:2025-05-13
  • Contact: *E-mail: yubo@henau.edu.cn; E-mail: guozhangkang@henau.edu.cn
  • Supported by:
    National Natural Science Foundation of China(32401739);Shennong Seed Industry Laboratory “First-class Project”(SN01-2022-01)

摘要:

挖掘小麦氮高效的种质和基因资源, 揭示其分子机制和遗传效应, 是当前小麦氮效率研究的重要内容和目标。本研究以255份不同小麦品种组成的自然群体为试验材料, 对每个品种1叶1心幼苗分别进行低氮(low nitrogen, LN, 0.05 mmol L-1 NO3-)和充足供氮(sufficient nitrogen, SN, 1.00 mmol L-1 NO3-) 2个水培条件处理, 培养28 d后, 在低氮和充足供氮处理下测定17个表型指标, 通过对55K SNP芯片进行质控过滤筛选出38,215个高质量单核苷酸多态性(single nucleotide polymorphism, SNPs)位点,结合FarmCPU、MLM以及MLM+Q+K模型进行全基因组关联分析(genome-wide association study, GWAS)。该群体在LN和SN水平下17个表型指标比值(LS, LN/SN)的频率分布均呈正态分布, 对17个LS指标进行全基因组关联分析, 共检测到1161个显著位点(P≤0.001), 发现有103个标记至少在2个模型中同时被检测到, 有8个SNP至少关联了4个性状, 其中AX-110548993 (3B)和AX-111802919 (4D)为新位点。2个新位点上下游5 Mb范围区间内含有267个候选基因, 其中位于4D染色体上的SNP AX-111802919包括3个直接参与或调控氮吸收转运的候选基因: TraesCS4D02G361500编码硝酸盐转运蛋白(NRT1.1), TraesCS4D02G362100编码锌指蛋白CONSTANS-LIKE 1, TraesCS4D02G362800编码1个GATA转录因子蛋白。

关键词: 小麦, 氮效率, 全基因组关联分析, 候选基因, 挖掘

Abstract:

To explore wheat germ plasm and genetic resources associated with high nitrogen use efficiency, and to elucidate their molecular mechanisms and genetic effects, are key objectives of current research on nitrogen efficiency in wheat. In this study, a natural population comprising 255 wheat varieties was used as experimental material. Seedlings at the one-leaf stage were subjected to hydroponic treatments under low nitrogen (LN, 0.05 mmol L-1 NO3-) and sufficient nitrogen (SN, 1.00 mmol L-1 NO3-) conditions. After 28 days, seventeen phenotypic traits were measured under both nitrogen treatments. A total of 38,215 high-quality single nucleotide polymorphism (SNP) sites were obtained through quality control filtering of the 55K SNP chip data. Genome-wide association studies (GWAS) were conducted using the FarmCPU, MLM, and MLM+Q+K models. The distribution of LN/SN ratios for the 17 traits (denoted as LS) followed a normal distribution across the population. GWAS identified 1161 significant loci (P ≤ 0.001) associated with the 17 LS traits, of which 103 SNPs were detected in at least two models. Notably, eight SNPs were associated with at least four traits, and two novel loci—AX-110548993 (on chromosome 3B) and AX-111802919 (on chromosome 4D)—were identified. Within a 5 Mb window upstream and downstream of these novel SNPs, 267 candidate genes were predicted. Specifically, SNP AX-111802919 on chromosome 4D includes three candidate genes directly involved in or regulating nitrogen uptake and transport: TraesCS4D02G361500, TraesCS4D02G362100, and TraesCS4D02G362800, which encode a nitrate transporter (NRT1.1), a zinc finger protein CONSTANS-LIKE 1, and a GATA transcription factor, respectively.

Key words: Triticum aestivum L., nitrogen efficiency, genome-wide association study, candidate gene, mining

附表1

255份小麦供试品种信息"

名称
Name
选育单位
Breeding unit
审定时间
Approval time
来源
Sources
皖麦38 Wanmai 38 安徽省宿州市农业科学院Anhui Academy of Agricultural Sciences 1997 中国安徽
Anhui, China
皖麦19 Wanmai 19 安徽省宿州市农业科学院Anhui Academy of Agricultural Sciences 1994 中国安徽
Anhui, China
皖麦50 Wanmai 50 安徽省宿州市农业科学院Anhui Academy of Agricultural Sciences 2004 中国安徽
Anhui, China
皖麦53 Wanmai 53 安徽省宿州市农业科学院 Anhui Academy of Agricultural sciences 2005 中国安徽
Anhui, China
皖麦52 Wanmai 52 安徽省宿州市种子公司Suzhou Seed Company in Anhui province 2004 中国安徽
Anhui, China
华成3366 Huacheng 3366 安徽省宿州市种子公司 Suzhou Seed Company in Anhui province 2013 中国安徽
Anhui, China
京411 Jing 411 北京市种子管理站Beijing Seed Management Station 1993 中国北京
Beijing, China
京9428 Jing 9428 北京市种子管理站Beijing Seed Management Station 2000 中国北京
Beijing, China
京冬12 Jingdong 12 北京杂交小麦工程技术研究中心
Beijing Hybrid wheat Engineering Technology Research Center
2004 中国北京
Beijing, China
京冬17 Jingdong 17 北京杂交小麦工程技术研究中心
Beijing Hybrid wheat Engineering Technology Research Center
2007 中国北京
Beijing, China
京冬22 Jingdong 22 北京杂交小麦工程技术研究中心
Beijing Hybrid wheat Engineering Technology Research Center
2007 中国北京
Beijing, China
京花9号 Jinghua 9 北京杂交小麦工程技术研究中心
Beijing Hybrid wheat Engineering Technology Research Center
2007 中国北京
Beijing, China
科农199 Kenong 199 中国科学院遗传与发育生物学研究所
Institute of Genetics and Developmental Biology, Chinese Academy of Sciences
2006 中国北京
Beijing, China
中育6号 Zhongyu 6 中国农业科学院棉花研究所
Institute of Cotton Research, Chinese Academy of Agricultural Sciences
2001 中国北京
Beijing, China
京冬8 Jingdong 8 中国农业科学院作物科学研究所
Institute of Crop Sciences, Chinese Academy of Agricultural Sciences
1996 中国北京
Beijing, China
轮选103 Lunxuan 103 中国农业科学院作物科学研究所
Institute of Crop Sciences, Chinese Academy of Agricultural Sciences
2015 中国北京
Beijing, China
轮选61 Lunxuan 61 中国农业科学院作物科学研究所
Institute of Crop Sciences, Chinese Academy of Agricultural Sciences
2009 中国北京
Beijing, China
轮选66 Lunxuan 66 中国农业科学院作物科学研究所
Institute of Crop Sciences, Chinese Academy of Agricultural Sciences
2018 中国北京
Beijing, China
轮选987 Lunxuan 987 中国农业科学院作物科学研究所
Institute of Crop Sciences, Chinese Academy of Agricultural Sciences
2003 中国北京
Beijing, China
中麦175 Zhongmai 175 中国农业科学院作物科学研究所
Institute of Crop Sciences, Chinese Academy of Agricultural Sciences
2012 中国北京
Beijing, China
中麦415 Zhongmai 415 中国农业科学院作物科学研究所
Institute of Crop Sciences, Chinese Academy of Agricultural Sciences
2010 中国北京
Beijing, China
中麦9号 Zhongmai 9 中国农业科学院作物科学研究所
Institute of Crop Sciences, Chinese Academy of Agricultural Sciences
1998 中国北京
Beijing, China
中麦8号 Zhongmai 8 中国农业科学院作物科学研究所
Institute of Crop Sciences, Chinese Academy of Agricultural Sciences
2016 中国北京
Beijing, China
中麦895 Zhongmai 895 中国农业科学院作物科学研究所
Institute of Crop Sciences, Chinese Academy of Agricultural Sciences
2012 中国北京
Beijing, China
保麦10 Baomai 10 保定市农业科学研究所 Baoding Agricultural Science Research Institute 2008 中国河北
Hebei, China
河农326 Henong 326 河北农业大学 Hebei Agricultural University 1997 中国河北
Hebei, China
河农6049 Henong 6049 河北农业大学 Hebei Agricultural University 2009 中国河北
Hebei, China
河农822 Henong 822 河北农业大学 Hebei Agricultural University 2004 中国河北
Hebei, China
藁2018 Gao 2018 河北省藁城市农业科学研究所
Agricultural Science Research Institute of Gaocheng, Hebei Province
2008 中国河北
Hebei, China
藁城8901 Gao 8901 河北省藁城市农业科学研究所
Agricultural Science Research Institute of Gaocheng, Hebei Province
1995 中国河北
Hebei, China
藁城9415 Gao 9415 河北省藁城市农业科学研究所
Agricultural Science Research Institute of Gaocheng, Hebei Province
2003 中国河北
Hebei, China
邯4564 Han 4564 河北省邯郸市农业科学院
Academy of Agricultural Sciences of Handan, Hebei Province
2006 中国河北
Hebei, China
邯4589 Han 4589 河北省邯郸市农业科学院
Academy of Agricultural Sciences of Handan, Hebei Province
2001 中国河北
Hebei, China
邯5316 Han 5316 河北省邯郸市农业科学院
Academy of Agricultural Sciences of Handan, Hebei Province
2000 中国河北
Hebei, China
邯6172 Han 6172 河北省邯郸市农业科学院
Academy of Agricultural Sciences of Handan, Hebei Province
2003 中国河北
Hebei, China
邯麦11 Hanmai 11 河北省邯郸市农业科学院
Academy of Agricultural Sciences of Handan, Hebei Province
2007 中国河北
Hebei, China
邯麦13 Hanmai 13 河北省邯郸市农业科学院
Academy of Agricultural Sciences of Handan, Hebei Province
2009 中国河北
Hebei, China
新冬20号 Xindong 20 河北省农林科学院Hebei Academy of Agriculture and Forestry 1995 中国河北
Hebei, China
衡4399 Heng 4399 河北省农林科学院旱作农业研究所
Institute of Dry Farming, Academy of Agriculture and Forestry Sciences of Hebei Province
2008 中国河北
Hebei, China
衡5229 Heng 5229 河北省农林科学院旱作农业研究所
Institute of Dry Farming, Academy of Agriculture and Forestry Sciences of Hebei Province
2004 中国河北
Hebei, China
冀5265 Ji 5265 河北省农林科学院粮油作物研究所
Institute of Cereal and Oil crops, Hebei Academy of Agriculture and Forestry Sciences
2009 中国河北
Hebei, China
金禾9123 Jinhe 9123 河北省农林科学院遗传生理研究所
Institute of Genetics and Physiology, Hebei Academy of Agriculture and Forestry Sciences
2012 中国河北
Hebei, China
冀麦26 Jimai 26 河北省石家庄市农业科学研究院
Academy of Agricultural Sciences of Shijiazhuang, Hebei Province
1991 中国河北
Hebei, China
冀麦30Jimai 30 河北省石家庄市农业科学研究院
Academy of Agricultural Sciences of Shijiazhuang, Hebei Province
1992 中国河北
Hebei, China
冀麦38 Jimai 38 河北省石家庄市农业科学研究院
Academy of Agricultural Sciences of Shijiazhuang, Hebei Province
1996 中国河北
Hebei, China
石4185 Shi 4185 河北省石家庄市农业科学研究院
Academy of Agricultural Sciences of Shijiazhuang, Hebei Province
1997 中国河北
Hebei, China
石7221 Shi 7221 河北省石家庄市农业科学研究院
Academy of Agricultural Sciences of Shijiazhuang, Hebei Province
2005 中国河北
Hebei, China
石家庄8号 Shijiazhuang 8 河北省石家庄市农业科学研究院
Academy of Agricultural Sciences of Shijiazhuang, Hebei Province
2001 中国河北
Hebei, China
石麦14号 Shimai 14 河北省石家庄市农业科学研究院
Academy of Agricultural Sciences of Shijiazhuang, Hebei Province
2004 中国河北
Hebei, China
邢麦4号 Xingmai 4 河北省邢台市农业科学院
Academy of Agricultural Sciences of Xingtai, Hebei Province
2007 中国河北
Hebei, China
师栾02-1 Shiluan 02-1 河北师范大学Hebei Normal University 2007 中国河北
Hebei, China
石麦15号Shimai 15 石家庄市农林科学研究院
Shijiazhuang Academy of Agriculture and Forestry Sciences
2009 中国河北
Hebei, China
石优20 Shiyou 20 石家庄市农林科学研究院
Shijiazhuang Academy of Agriculture and Forestry Sciences
2011 中国河北
Hebei, China
石新733 Shixin 733 石家庄市小麦新品种新技术研究所
Shijiazhuang Wheat New Varieties New Technology Institute
2001 中国河北
Hebei, China
石新828 Shixin 828 石家庄市小麦新品种新技术研究所
Shijiazhuang Wheat New Varieties New Technology Institute
2013 中国河北
Hebei, China
高优503 Gaoyou 503 中国科学院石家庄农业现代化研究所
Shijiazhuang Institute of Agricultural Modernization, Chinese Academy of Sciences
2001 中国河北
Hebei, China
农大3432 Nongda 3432 中国农大农学与生物技术学院
College of Agronomy and Biotechnology, China Agricultural University
2005 中国河北
Hebei, China
衡观35 Hengguan 35 河北省农林科学院旱作农业研究所
Institute of dry farming, Academy of Agriculture and Forestry Sciences of Hebei Province
2004 中国河北
Hebei, China
怀川916 Huaichuan 916 河南怀川种业有限责任公司
Henan Huaichuan Seed Industry Limited Liability Company
2001 中国河南
Henan, China
豫教5号 Yujiao 5 河南教育学院 Henan Institute of Education 2011 中国河南
Henan, China
金丰3号 Jinfeng 3 河南科技大学农学院
College of Agriculture, Henan University of Science and Technology
2005 中国河南
Henan, China
百农160 Bainong 160 河南科技学院 Henan University of Science and Technology 2007 中国河南
Henan, China
豫麦52号 Yumai 52 河南农业大学 Henan Agricultural University 1998 中国河南
Henan, China
豫农202 Yunong 202 河南农业大学 Henan Agricultural University 2007 中国河南
Henan, China
豫农416 Yunong 416 河南农业大学 Henan Agricultural University 2009 中国河南
Henan, China
豫农949 Yunong 949 河南农业大学 Henan Agricultural University 2005 中国河南
Henan, China
豫农982 Yunong 982 河南农业大学 Henan Agricultural University 2009 中国河南
Henan, China
平安8号 Pingan 8 河南平安种业有限公司 Henan Pingan Seed Industry Limited Company 2012 中国河南
Henan, China
豫麦49-198 Yumai 49-198 河南平安种业有限公司 Henan Pingan Seed Industry Limited Company 2005 中国河南
Henan, China
百3217 Bai 3217 河南省百泉农校 Henan Province Baiquan Agricultural School 1984 中国河南
Henan, China
豫麦2号 Yumai 2 河南省宝丰县农业科学研究所
Agricultural Science Institute of Baofeng County, Henan Province
1984 中国河南
Henan, China
博农7023 Bonong 7023 河南省博爱农场 Boai farm in Henan Province 1981 中国河南
Henan, China
泛麦5号 Fanmai 5 河南省地神种业农业科学研究院
Henan Province Dishen Seed Industry Agricultural Science Research Institute
2004 中国河南
Henan, China
泛麦8号 Fanmai 8 河南省地神种业农业科学研究院
Henan Province Dishen Seed Industry Agricultural Science Research Institute
2008 中国河南
Henan, China
富麦2008 Fumai 2008 河南省科学院同位素研究所
Institute of Isotopes, Henan Academy of Sciences
2006 中国河南
Henan, China
漯麦4号 Luomai 4 河南省漯河市农业科学院
Luohe Academy of Agricultural Sciences, Henan Province
2003 中国河南
Henan, China
漯麦8号 Luomai 8 河南省漯河市农业科学院
Luohe Academy of Agricultural Sciences, Henan Province
2007 中国河南
Henan, China
豫麦70-36 Yumai 70-36 河南省内乡县农业科学研究所
Henan Neixiang County agricultural science research Institute
2004 中国河南
Henan, China
豫麦70号 Yumai 70 河南省内乡县农业科学研究所
Henan Neixiang County Agricultural Science Research Institute
2003 中国河南
Henan, China
郑麦002 Zhengmai 002 河南省农业科学院 Henan Academy of Agricultural Sciences 2003 中国河南
Henan, China
豫麦13号 Yumai 13 河南省农业科学院 Henan Academy of Agricultural Sciences 1993 中国河南 Henan, China
豫麦29号 Yumai 29 河南省农业科学院 Henan Academy of Agricultural Sciences 2014 中国河南 Henan, China
豫麦47号 Yumai 47 河南省农业科学院 Henan Academy of Agricultural Sciences 1997 中国河南 Henan, China
郑丰9962
Zhengfeng 9962
河南省农业科学院 Henan Academy of Agricultural Sciences 2009 中国河南
Henan, China
郑麦004 Zhengmai 004 河南省农业科学院 Henan Academy of Agricultural Sciences 2004 中国河南
Henan, China
郑麦366 Zhengmai 366 河南省农业科学院 Henan Academy of Agricultural Sciences 2005 中国河南
Henan, China
郑麦7698 Zhengmai 7698 河南省农业科学院 Henan Academy of Agricultural Sciences 2012 中国河南
Henan, China
郑麦9023 Zhengmai 9023 河南省农业科学院 Henan Academy of Agricultural Sciences 2001 中国河南
Henan, China
郑麦9962 Zhengmai 9962 河南省农业科学院 Henan Academy of Agricultural Sciences 2009 中国河南
Henan, China
花培3号 Huapei 3 河南省农业科学院生物技术研究所
Biotechnology Institute, Henan Academy of Agricultural Sciences
2006 中国河南
Henan, China
花培5号 Huapei 5 河南省农业科学院生物技术研究所
Biotechnology Institute, Henan Academy of Agricultural Sciences
2006 中国河南
Henan, China
花培8号 Huapei 8 河南省农业科学院生物技术研究所
Biotechnology Institute, Henan Academy of Agricultural Sciences
2009 中国河南
Henan, China
豫保1号 Yubao 1 河南省农业科学院植物保护研究所
Institute of Plant Protection, Henan Academy of Agricultural Sciences
2008 中国河南
Henan, China
濮麦9号 Pumai 9 河南省濮阳农业科学研究所
Henan Puyang Institute of Agricultural Sciences
2005 中国河南
Henan, China
丰德存麦1号Fengdecunmai 1 河南省天存小麦改良技术研究所
Henan Tiancun Wheat improvement Technology Research Institute
2011 中国河南
Henan, China
众麦2号 Zhongmai 2 河南省天宁种业有限公司 Heinan Tianning Seed Industry Limited Company 2006 中国河南
Henan, China
温9519 Wen 9519 河南省温县农科所 Wenxian Agricultural Science Institute of Henan Province 2004 中国河南
Henan, China
温9629 Wen 9629 河南省温县农科所 Wenxian Agricultural Science Institute of Henan Province 2004 中国河南
Henan, China
豫麦49号 Yumai 49 河南省温县祥云镇农技站 Henan province Wenxian Xiangyun town agricultural technology station 2000 中国河南
Henan, China
豫麦58号 Yumai 58 河南省温县种子公司 Henan Wenxian seed company 2001 中国河南
Henan, China
豫麦54号 Yumai 54 河南省新乡百泉技术师院小麦育种中心
Henan Province Xinxiang Baiquan Technical Teachers Institute Wheat Breeding Center
1998 中国河南
Henan, China
百泉3039 Baiquan 3039 河南省新乡市农业科学院 Xinxiang Academy of Agricultural Sciences, Henan Province 1981 中国河南
Henan, China
新麦11 Xinmai 11 河南省新乡市农业科学院 Xinxiang Academy of Agricultural Sciences, Henan Province 2001 中国河南
Henan, China
新麦18 Xinmai 18 河南省新乡市农业科学院 Xinxiang Academy of Agricultural Sciences, Henan Province 2004 中国河南
Henan, China
新麦19 Xinmai 19 河南省新乡市农业科学院 Xinxiang Academy of Agricultural Sciences, Henan Province 2006 中国河南
Henan, China
新麦21 Xinmai 21 河南省新乡市农业科学院 Xinxiang Academy of Agricultural Sciences, Henan Province 2009 中国河南
Henan, China
新麦9号 Xinmai 9 河南省新乡市农业科学院 Xinxiang Academy of Agricultural Sciences, Henan Province 2000 中国河南
Henan, China
许科1号 Xuke 1 河南省许科种业有限公司 Heinan Xuke Seed Industry Limited Company 2007 中国河南
Henan, China
许科316 Xuke 316 河南省许科种业有限公司 Heinan Xuke Seed Industry Limited Company 2011 中国河南
Henan, China
许科718 Xuke 718 河南省许科种业有限公司 Heinan Xuke Seed Industry Limited Company 2012 中国河南
Henan, China
偃高1号 Yangao 1 河南省偃师市高龙镇农技站
Henan Yanshi City Gaolong Town Agricultural Technology Station
2002 中国河南
Henan, China
亿麦6号 Yimai 6 河南省亿源种业有限公司 Heinan Yiyuan Seed Industry Limited Company 2011 中国河南
Henan, China
豫麦10号 Yumai 10 河南省豫西农业专科学校 Henan Yuxi Agricultural College 1991 中国河南
Henan, China
偃展1号 Yanzhan 1 河南省豫西农作物品种展览中心
Henan Yuxi Crop Variety Exhibition Center
2000 中国河南
Henan, China
偃展4110 Yanzhan 4110 河南省豫西农作物品种展览中心
Henan Yuxi crop Variety Exhibition Center
2003 中国河南
Henan, China
豫麦18号 Yumai 18 河南省豫西农作物品种展览中心
Henan Yuxi Crop Variety Exhibition Center
1995 中国河南
Henan, China
豫麦21号 Yumai 21 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
1994 中国河南
Henan, China
周8425B Zhou 8425B 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
1988 中国河南
Henan, China
周麦11 Zhoumai 11 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
1998 中国河南
Henan, China
周麦16 Zhoumai 16 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
2003 中国河南
Henan, China
周麦17 Zhoumai 17 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
2004 中国河南
Henan, China
周麦18 Zhoumai 18 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
2005 中国河南
Henan, China
周麦19 Zhoumai 19 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
2005 中国河南
Henan, China
周麦20 Zhoumai 20 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
2003 中国河南
Henan, China
周麦22 Zhoumai 22 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
2007 中国河南
Henan, China
周麦23 Zhoumai 23 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
2008 中国河南
Henan, China
周麦24 Zhoumai 24 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
2009 中国河南
Henan, China
周麦25 Zhoumai 25 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
2011 中国河南
Henan, China
周麦26 Zhoumai 26 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
2012 中国河南
Henan, China
周麦27 Zhoumai 27 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
2011 中国河南
Henan, China
周麦32 Zhoumai 32 河南省周口市农业科学院
Zhoukou Academy of Agricultural Sciences, Henan Province
2018 中国河南
Henan, China
众麦1号 Zhongmai 1 河南睢县农业科学研究所
Henan Suixian Agricultural Science Research Institute
2004 中国河南
Henan, China
天民198 Tianmin 198 河南天民种业有限公司 Henan Tianmin Seed Industry Limited Company 2014 中国河南
Henan, China
开麦18号 Kaimai 18 开封市农林科学研究院
Kaifeng Academy of Agriculture and Forestry Sciences
2004 中国河南
Henan, China
开麦20号 Kaimai 20 开封市农林科学研究院
Kaifeng Academy of Agriculture and Forestry Sciences
2011 中国河南
Henan, China
开麦21号 Kaimai 21 开封市农林科学研究院
Kaifeng Academy of Agriculture and Forestry Sciences
2011 中国河南
Henan, China
洛旱11 Luohan 11 洛阳市农林科学院
Luoyang Academy of Agriculture and Forestry Sciences
2008 中国河南
Henan, China
洛旱2号 Luohan 2 洛阳市农林科学院
Luoyang Academy of Agriculture and Forestry Sciences
2003 中国河南
Henan, China
洛旱6号 Luohan 6 洛阳市农林科学院
Luoyang Academy of Agriculture and Forestry Sciences
2006 中国河南
Henan, China
洛麦21 Luomai 21 洛阳市农林科学院
Luoyang Academy of Agriculture and Forestry Sciences
2009 中国河南
Henan, China
洛麦22 Luomai 22 洛阳市农林科学院
Luoyang Academy of Agriculture and Forestry Sciences
2008 中国河南
Henan, China
洛麦23 Luomai 23 洛阳市农林科学院
Luoyang Academy of Agriculture and Forestry Sciences
2009 中国河南
Henan, China
洛麦24 Luomai 24 洛阳市农林科学院
Luoyang Academy of Agriculture and Forestry Sciences
2013 中国河南
Henan, China
洛麦26 Luomai 26 洛阳市农林科学院
Luoyang Academy of Agriculture and Forestry Sciences
2018 中国河南
Henan, China
太学7号 Taixue 7 洛阳太学农作物研究所 Luoyang Taixue Crop Research Institute 2011 中国河南
Henan, China
南阳75-6 Nanyang 75-6 南阳农业科学院 Nanyang Academy of Agricultural Sciences 1976 中国河南
Henan, China
平麦998 Pingmai 998 平顶山市农业科学院 Pingdingshan Academy of Agricultural Sciences 2008 中国河南
Henan, China
汝州0319 Ruzhou 0319 汝州市农业科学研究所
Ruzhou Agricultural Science Research Institute
2009 中国河南
Henan, China
舜麦1718 Shunmai 1718 山西省农业科学院棉花研究所
Cotton Research Institute, Shanxi Academy of Agricultural Sciences
2011 中国河南
Henan, China
丰舞981 Fengwu 981 舞阳县种子公司 Wuyang County seed company 2004 中国河南
Henan, China
矮丰3号 Aifeng 3 西北农林科技大学Northwest Agriculture and Forestry University 1979 中国河南
Henan, China
丰产3号 Fengchan 3 西北农林科技大学 Northwest Agriculture and Forestry University 1964 中国河南
Henan, China
小偃22 Xiaoyan 22 西北农林科技大学 Northwest Agriculture and Forestry University 2003 中国河南
Henan, China
众麦998 Zhongmai 998 新安县科协郭根
Xin’an County Science and Technology Association Guo Gen
2008 中国河南
Henan, China
新原958 Xinyuan 958 新乡县原种一场 Xinxiang County Original Plant a Field 2005 中国河南
Henan, China
豫麦7号Yumai 7 偃师县农科所 Yanshi County Agricultural Science Institute 1985 中国河南
Henan, China
郑农16 Zhengnong 16 郑州市农林科学研究所
Zhengzhou Institute of Agriculture and Forestry Science
2004 中国河南
Henan, China
郑农17 Zhengnong 17 郑州市农林科学研究所 Zhengzhou Institute of Agriculture and Forestry Science 2004 中国河南
Henan, China
豫麦34号 Yumai 34 郑州市农林科学研究所 Zhengzhou Institute of Agriculture and Forestry Science 1998 中国河南
Henan, China
郑育麦9987
Zhengyumai 9987
郑州市友邦农作物新品种研究所
Zhengzhou Youbang Institute of New crop varieties
2009 中国河南
Henan, China
小偃81 Xiaoyan 81 中国科学院遗传与发育生物学研究所
Institute of Genetics and Developmental Biology, Chinese Academy of Sciences
2005 中国河南
Henan, China
中育10号 Zhongyu 10 中国农业科学院棉花研究所
Institute of Cotton Research, Chinese Academy of Agricultural Sciences
2007 中国河南
Henan, China
04中36 04 Zhong 36 中国农业科学院棉花研究所
Institute of Cotton Research, Chinese Academy of Agricultural Sciences
2006 中国河南
Henan, China
中育12号 Zhongyu 12 中国农业科学院棉花研究所
Institute of Cotton Research, Chinese Academy of Agricultural Sciences
2008 中国河南
Henan, China
中育9号 Zhongyu 9 中国农业科学院棉花研究所
Institute of Cotton Research, Chinese Academy of Agricultural Sciences
2004 中国河南
Henan, China
中麦875 Zhongmai 875 中国农业科学院作物科学研究所
Institute of Crop Science, Chinese Academy of Agricultural Sciences
2014 中国河南
Henan, China
中育9398 Zhongyu 9398 中棉种业科技股份有限公司 China Cotton Seed Technology Stock Limited Company 2012 中国河南
Henan, China
驻麦4号 Zhumai 4 驻马店市农业科学院 Zhumadian Academy of Agricultural Sciences 2004 中国河南
Henan, China
垦九10号 Kenjiu 10 黑龙江省农垦总局九三科学研究所
Heilongjiang Province General Bureau of Land Reclamation 93 Scientific Research Institute
2003 中国黑龙江 Heilongjiang, China
新克旱9 Xinkehan 9 黑龙江省农业科学院(克山分院)
Heilongjiang Academy of Agricultural Sciences (Keshan Branch)
1988 中国黑龙江 Heilongjiang, China
保麦2号 Baomai 2 江苏保丰集团公司 Jiangsu Baofeng Group Company 2012 中国江苏 Jiangsu, China
扬麦12号 Yangmai 12 江苏里下河地区农业科学研究所
Jiangsu Lixiahe Area Agricultural Science Research Institute
2001 中国江苏 Jiangsu, China
扬麦15号 Yangmai 15 江苏里下河地区农业科学研究所
Jiangsu Lixiahe Area Agricultural Science Research Institute
2005 中国江苏 Jiangsu, China
扬麦16号 Yangmai 16 江苏里下河地区农业科学研究所
Jiangsu Lixiahe Area Agricultural Science Research Institute
2004 中国江苏 Jiangsu, China
淮麦18号 Huaimai 18 江苏省淮阴市农业科学研究所
Huaiyin Institute of Agricultural Sciences, Jiangsu Province
2001 中国江苏 Jiangsu, China
连麦2号 Lianmai 2 江苏省连云港市农业科学院
Lianyungang Academy of Agricultural Sciences, Jiangsu Province
2005 中国江苏 Jiangsu, China
宁麦13 Ningmai 13 江苏省农业科学院粮食作物研究所
Food Crop Research Institute, Jiangsu Academy of Agricultural Sciences
2006 中国江苏 Jiangsu, China
徐州24 Xuzhou 24 江苏省徐州市农业科学院Xuzhou Academy of Agricultural Sciences, Jiangsu Province 1994 中国江苏 Jiangsu, China
徐州25 Xuzhou 25 江苏省徐州市农业科学院
Xuzhou Academy of Agricultural Sciences, Jiangsu Province
2000 中国江苏 Jiangsu, China
淮麦20号 Huaimai 20 江苏徐淮地区淮阴农业科学研究所
Jiangsu Xuhuai area Huaiyin Agricultural science research Institute
2002 中国江苏 Jiangsu, China
淮麦21号 Huaimai 21 江苏徐淮地区淮阴农业科学研究所
Jiangsu Xuhuai area Huaiyin Agricultural science research Institute
2008 中国江苏 Jiangsu, China
淮麦22号 Huaimai 22 江苏徐淮地区淮阴农业科学研究所
Jiangsu Xuhuai area Huaiyin Agricultural science research Institute
2007 中国江苏 Jiangsu, China
淮麦28号 Huaimai 28 江苏徐淮地区淮阴农业科学研究所
Jiangsu Xuhuai area Huaiyin Agricultural science research Institute
2009 中国江苏 Jiangsu, China
淮麦29号 Huaimai 29 江苏徐淮地区淮阴农业科学研究所
Jiangsu Xuhuai area Huaiyin Agricultural science research Institute
2009 中国江苏 Jiangsu, China
徐麦30 Xumai 30 江苏徐淮地区淮阴农业科学研究所
Jiangsu Xuhuai area Huaiyin Agricultural science research Institute
2007 中国江苏 Jiangsu, China
徐麦856 Xumai 856 江苏徐淮地区淮阴农业科学研究所
Jiangsu Xuhuai area Huaiyin Agricultural science research Institute
2006 中国江苏 Jiangsu, China
92R137 南京农业大学 Nanjing Agricultural University 1998 中国江苏 Jiangsu, China
宁春4 Ningchun 4 宁夏永宁县良种繁殖场
Yongning county, Ningxia improved seed breeding farm
1981 中国宁夏 Ningxia, China
良星99 Liangxing 99 山东德州市良星种子研究所
Shandong Dezhou Liangxing seed Research Institute
2006 中国山东 Shandong, China
鲁麦15号 Lumai 15 山东农业大学 Shandong Agricultural University 1996 中国山东 Shandong, China
鲁垦麦9号 Lukenmai 9 山东仁和种业有限责任公司
Shandong Renhe seed industry limited liability company
2010 中国山东 Shandong, China
鲁原502 Luyuan 502 山东省农业科学院原子能农业应用研究所
Atomic Energy Agricultural Application Research Institute, Shandong Academy of Agricultural Sciences
2001 中国山东 Shandong, China
山农20 Shannong 20 山东省农业科学院原子能农业应用研究所
Atomic Energy Agricultural Application Research Institute, Shandong Academy of Agricultural Sciences
2011 中国山东 Shandong, China
济麦19 Jimai 19 山东省农业科学院作物研究所
Crop Research Institute, Shandong Academy of Agricultural Sciences
2003 中国山东 Shandong, China
济麦20 Jimai 20 山东省农业科学院作物研究所
Crop Research Institute, Shandong Academy of Agricultural Sciences
2004 中国山东Shandong, China
济麦21 Jimai 21 山东省农业科学院作物研究所
Crop Research Institute, Shandong Academy of Agricultural Sciences
2004 中国山东 Shandong, China
济麦22 Jimai 22 山东省农业科学院作物研究所
Crop Research Institute, Shandong Academy of Agricultural Sciences
2006 中国山东 Shandong, China
济南13 Jinan 13 山东省农业科学院作物研究所
Crop Research Institute, Shandong Academy of Agricultural Sciences
1990 中国山东 Shandong, China
济南16 Jinan 16 山东省农业科学院作物研究所
Crop Research Institute, Shandong Academy of Agricultural Sciences
1998 中国山东 Shandong, China
济南17 Jinan 17 山东省农业科学院作物研究所
Crop Research Institute, Shandong Academy of Agricultural Sciences
1999 中国山东 Shandong, China
泰山21号 Taishan 21 山东省泰安市农业科学研究院
Tai’an Academy of Agricultural Sciences, Shandong Province
2003 中国山东 Shandong, China
泰山23号 Taishan 23 山东省泰安市农业科学研究院
Tai’an Academy of Agricultural Sciences, Shandong Province
2004 中国山东 Shandong, China
郯麦98 Tanmai 98 山东省郯城县种子公司
Tancheng County seed company, Shandong province
2009 中国山东Shandong, China
鲁麦21号 Lumai 21 山东省烟台市农业科学研究院
Yantai Academy of Agricultural Sciences, Shandong Province
1996 中国山东 Shandong, China
鲁麦14号 Lumai 14 山东省烟台市农业科学研究院
Yantai Academy of Agricultural Sciences, Shandong Province
1990 中国山东 Shandong, China
鲁麦7号 Lumai 7 山东省烟台市农业科学研究院
Yantai Academy of Agricultural Sciences, Shandong Province
1984 中国山东 Shandong, China
烟辐188 Yanfu 188 山东省烟台市农业科学研究院
Yantai Academy of Agricultural Sciences, Shandong Province
2002 中国山东 Shandong, China
烟农15号 Yannong 15 山东省烟台市农业科学研究院
Yantai Academy of Agricultural Sciences, Shandong Province
1982 中国山东 Shandong, China
烟农19号 Yannong 19 山东省烟台市农业科学研究院
Yantai Academy of Agricultural Sciences, Shandong Province
2001 中国山东 Shandong, China
烟农21号 Yannong 21 山东省烟台市农业科学研究院
Yantai Academy of Agricultural Sciences, Shandong Province
2004 中国山东 Shandong, China
烟农836 Yannong 836 山东省烟台市农业科学研究院
Yantai Academy of Agricultural Sciences, Shandong Province
2010 中国山东 Shandong, China
烟农999 Yannong 999 山东省烟台市农业科学研究院
Yantai Academy of Agricultural Sciences, Shandong Province
2016 中国山东 Shandong, China
蚰包麦 Youbaomai 山东省烟台市农业科学研究院
Yantai Academy of Agricultural Sciences, Shandong Province
1972 中国山东 Shandong, China
洲元9369 Zhouyuan 9369 山东洲元种业股份有限公司
Shandong Zhouyuan Seed Industry Stock Limited Company
2007 中国山东 Shandong, China
鲁麦23号 Lumai 23 胜利油田滨南马坊农场
Shengli oil field Binnan Ma Fang farm
1996 中国山东 Shandong, China
泰农18 Tainong 18 泰安市泰山区瑞丰作物育种研究所
Ruifeng Crop Breeding Institute, Taishan District, Tai’an City
2008 中国山东 Shandong, China
汶农14 Wennong 14 泰安市汶农种业有限责任公司
Tai’an Wennong seed industry limited liability company
2011 中国山东 Shandong, China
淄麦12号 Zimai 12 淄博市农业科学研究院 Zibo Academy of Agricultural Sciences 2001 中国山东 Shandong, China
晋麦47 Jinmai 47 山西省农科院棉花研究所
Institute of Cotton Research, Shanxi Academy of Agricultural Sciences
1995 中国山西
Shanxi, China
尧麦16 Yaomai 16 山西省农业科学院小麦研究所
Institute of Wheat Research, Shanxi Academy of Agricultural Sciences
2011 中国山西
Shanxi, China
陕229 Shaan 229 陕西省农业科学院 Shaanxi Academy of Agricultural Sciences 1996 中国陕西 Shaanxi, China
西安8号 Xi′an 8 陕西省西安市农业科学所
Xi’an Institute of Agricultural Sciences, Shaanxi Province
1985 中国陕西 Shaanxi, China
西农979 Xinong 979 西北农林科技大学 Northwest Agriculture & Forestry University 2005 中国陕西 Shaanxi, China
小偃54 Xiaoyan 54 西北农林科技大学 Northwest Agriculture & Forestry University 2000 中国陕西 Shaanxi, China
绵农4号 Miannong 4 绵阳经济专科学校 Mianyang Economics College 1993 中国四川 Sichuan, China
绵阳11号 Mianyang 11 绵阳市农业科学研究院 Mianyang Academy of Agricultural Sciences 1994 中国四川 Sichuan, China
绵阳19号 Mianyang 19 绵阳市农业科学研究院 Mianyang Academy of Agricultural Sciences 1991 中国四川 Sichuan, China
绵阳20号 Mianyang 20 绵阳市农业科学研究院 Mianyang Academy of Agricultural Sciences 1990 中国四川 Sichuan, China
绵阳26号 Mianyang 26 绵阳市农业科学研究院 Mianyang Academy of Agricultural Sciences 1998 中国四川 Sichuan, China
川农16号 Chuannong 16 四川农业大学 Sichuan Agricultural University 2003 中国四川 Sichuan, China
繁6 Fan 6 四川农业大学 Sichuan Agricultural University 1978 中国四川 Sichuan, China
内麦8号 Neimai 8 四川省内江市农业科学院 Neijiang Academy of Agricultural Sciences, Sichuan Province 2003 中国四川 Sichuan, China
川麦107 Chuanmai 107 四川省农业科学院作物研究所
Crop Research Institute, Sichuan Academy of Agricultural Sciences
2000 中国四川 Sichuan, China
川麦22 Chuanmai 22 四川省农业科学院作物研究所
Crop Research Institute, Sichuan Academy of Agricultural Sciences
1989 中国四川 Sichuan, China
川麦24 Chuanmai 24 四川省农业科学院作物研究所
Crop Research Institute, Sichuan Academy of Agricultural Sciences
1994 中国四川 Sichuan, China
川麦28 Chuanmai 28 四川省农业科学院作物研究所
Crop Research Institute, Sichuan Academy of Agricultural Sciences
1997 中国四川 Sichuan, China
川麦42 Chuanmai 42 四川省农业科学院作物研究所
Crop Research Institute, Sichuan Academy of Agricultural Sciences
2003 中国四川 Sichuan, China
川麦46 Chuanmai 46 四川省农业科学院作物研究所
Crop Research Institute, Sichuan Academy of Agricultural Sciences
2005 中国四川Sichuan, China
川麦47 Chuanmai 47 四川省农业科学院作物研究所
Crop Research Institute, Sichuan Academy of Agricultural Sciences
2005 中国四川Sichuan, China
良星66 Liangxing 66 山东良星种业有限公司
Shandong Liangxing Seed Industry Limited Company
2008 中国天津
Tianjin, China
阿勃 Abo 西藏自治区农业科学研究所
Tibet Autonomous Region Institute of Agricultural Sciences
1985 中国西藏
Xizang, China
扬麦13号 Yangmai 13 江苏省里下河地区农业科学研究所
Lixiahe Area Agricultural Science Research Institute, Jiangsu Province
2002 中国江苏
Jiangsu, China
扬麦158 Yangmai 158 江苏省里下河地区农业科学研究所
Lixiahe Area Agricultural Science Research Institute, Jiangsu Province
1997 中国江苏
Jiangsu, China
轮选126 Lunxuan 126 中国农业科学院作物科学研究所
Institute of Crop Science, Chinese Academy of Agricultural Sciences
未知 Unknown 中国北京
Beijing, China
轮选136 Lunxuan 136 中国农业科学院作物科学研究所
Institute of Crop Science, Chinese Academy of Agricultural Sciences
未知 Unknown 中国北京
Beijing, China
轮选154Lunxuan 154 中国农业科学院作物科学研究所
Institute of Crop Science, Chinese Academy of Agricultural Sciences
未知 Unknown 中国北京
Beijing, China
轮选199Lunxuan 199 中国农业科学院作物科学研究所
Institute of Crop Science, Chinese Academy of Agricultural Sciences
未知 Unknown 中国北京
Beijing, China
欧柔Orofen 未知 Unknown 未知 Unknown 智利Chile
鲁98421 Lu98421 未知 Unknown 未知 Unknown 未知 Unknown
安农1008 Annong 1008 未知 Unknown 未知 Unknown 未知 Unknown
马场2号 Machang 2 未知 Unknown 未知 Unknown 未知 Unknown
宿3596 Su 3596 未知 Unknown 未知 Unknown 未知 Unknown
郑州3号 Zhengzhou 3 未知 Unknown 未知 Unknown 未知 Unknown
郑州761 Zhongzhou 761 未知 Unknown 未知 Unknown 未知 Unknown
中泛4 Zhongfan 4 未知 Unknown 未知 Unknown 未知 Unknown

表1

小麦苗期氮效率相关性状表型统计"

性状
Trait
充足供氮
Sufficient nitrogen
低氮
Low nitrogen
低氮/充足供氮
Low nitrogen/sufficient nitrogen
平均值
Mean
标准差
SD
变异系数
CV (%)
平均值
Mean
标准差
SD
变异系数
CV (%)
平均值
Mean
标准差
SD
变异系数
CV (%)
SFW (g plant-1) 2.420 a 0.815 33.654 0.718 b 0.242 33.724
RFW (g plant-1) 1.730 a 0.638 36.876 0.961 b 0.358 37.228
TFW (g plant-1) 4.150 a 1.401 33.763 1.679 b 0.571 34.003
SDW (g plant-1) 0.418 a 0.154 36.876 0.166 b 0.058 35.144
RDW (g plant-1) 0.130 a 0.043 33.347 0.090 b 0.044 48.847
TDW (g plant-1) 0.548 a 0.188 34.361 0.256 b 0.092 36.047
SNC (g plant-1) 21.913 a 4.952 22.596 10.882 b 1.778 16.341
RNC (g kg-1) 16.616 a 2.965 17.845 11.263 b 1.526 13.547
TNC (g kg-1) 20.532 a 3.836 18.684 10.974 b 1.394 12.703
SNA (g plant-1) 9.091 a 3.818 42.004 1.778 b 0.636 35.793
RNA (g plant-1) 2.137 a 0.777 36.366 1.004 b 0.493 49.098
TNA (g plant-1) 11.228 a 4.405 39.229 2.782 b 1.013 36.417
SNUE (g g-1 kg-1) 0.020 a 0.009 45.923 0.016 b 0.007 41.471
RNUE (g g-1 kg-1) 0.008 a 0.003 38.784 0.008 a 0.004 51.962
TNUE (g g-1 kg-1) 0.028 a 0.011 39.742 0.024 b 0.009 39.724
RSDW 0.328 b 0.107 32.642 0.550 a 0.171 31.124
RSNA 0.252 b 0.088 35.029 0.572 a 0.188 32.904
LS-SFW 0.328 0.161 49.092
LS-RFW 0.636 0.388 60.943
LS-TFW 0.450 0.229 50.899
LS-SDW 0.455 0.255 55.967
LS-RDW 0.761 0.437 57.397
LS-TDW 0.522 0.273 52.325
LS-SNC 0.516 0.124 23.991
LS-RNC 0.700 0.165 23.596
LS-TNC 0.550 0.110 19.959
LS-SNA 0.228 0.129 56.810
LS-RNA 0.522 0.303 58.018
LS-TNA 0.282 0.152 53.950
LS-SNUE 0.959 0.608 63.411
LS-RNUE 1.167 0.798 68.354
LS-TNUE 1.005 0.581 57.806
LS-RSDW 1.789 0.655 36.627
LS-RSNA 2.461 1.066 43.320

图1

255份小麦品种氮效率相关性状LS值的频率分布 缩写同表1。"

图2

255份小麦品种氮效率相关性状LS值的相关性分析 缩写同表1。*、**和***分别表示在0.05、0.01和0.001水平相关性显著。"

图3

SNP标记在小麦21条染色体上的分布"

表2

SNP标记密度分析"

染色体
Chromosome
SNP数
SNP
number
SNP覆盖距离
Coverage
distance of SNP (Mb)
标记密度
Density of
markers
(Mb marker−1)
染色体
Chromosome
SNP数
SNP
number
SNP覆盖距离
Coverage
distance of SNP (Mb)
标记密度
Density of
markers
(Mb marker−1)
1A 2060 593.28 0.29 4D 650 509.27 0.78
1B 2103 688.70 0.33 5A 2173 709.19 0.33
1D 1786 495.23 0.28 5B 2225 712.40 0.32
2A 2293 780.76 0.34 5D 1360 564.75 0.42
2B 1959 801.23 0.41 6A 2132 617.99 0.29
2D 1581 650.78 0.41 6B 2085 720.49 0.35
3A 1648 750.47 0.46 6D 1236 473.52 0.38
3B 2201 829.73 0.38 7A 2028 735.46 0.36
3D 1444 615.49 0.43 7B 1918 750.55 0.39
4A 1339 743.32 0.56 7D 1694 636.61 0.38
4B 2300 673.47 0.29 All 38,215 14,052.69 0.37

图4

255份供试小麦品种的群体结构及连锁不平衡分析 A: K取不同值时ΔK的对应分布; B: 群体结构, I~VIII表示群体分为8个亚群; C: 全基因组连锁不平衡(LD)衰减曲线。"

附图1

基于255份小麦品种氮效率相关性状LS值的全基因组关联分析(GWAS) 缩写同表1。曼哈顿图: 从里环到外环分别为各氮效率相关性状的FarmCPU、MLM以及MLM+Q+K模型。"

表3

小麦氮效率全基因组关联分析定位稳定SNP标记"

性状
Trait
标记
SNP
染色体
Chromosome
位置
Position (Mb)
模型
Model
P最大值
P max.
P最小值
P min.
贡献率
R2 (%)
LS-SFW AX-111660453 1A 574.476,865 FarmCPU, MLM, MLM+Q+K 1.80E-04 1.89E-05 8.077
AX-109551739 1A 574.486,675 FarmCPU, MLM, MLM+Q+K 1.80E-04 1.89E-05 8.077
AX-111802919 4D 508.503,834 FarmCPU, MLM+Q+K 8.65E-04 1.70E-04 8.400
LS-RDW AX-111660453 1A 574.476,865 FarmCPU, MLM, MLM+Q+K 8.41E-04 1.78E-04 6.553
AX-109551739 1A 574.486,675 FarmCPU, MLM, MLM+Q+K 8.41E-04 1.78E-04 6.553
LS-SDW AX-111660453 1A 574.476,865 FarmCPU, MLM, MLM+Q+K 1.62E-04 2.05E-05 8.051
AX-109551739 1A 574.486,675 FarmCPU, MLM, MLM+Q+K 1.62E-04 2.05E-05 8.051
AX-110920832 3B 44.975,458 FarmCPU, MLM 4.26E-04 9.27E-05 4.890
AX-110979247 3B 45.045,393 FarmCPU, MLM 3.95E-04 8.40E-05 5.050
AX-111518365 3B 45.289,556 FarmCPU, MLM 2.83E-04 5.43E-05 5.500
AX-109940356 3B 45.257,506 FarmCPU, MLM 6.19E-04 1.50E-04 4.670
AX-110548993 3B 65.160,500 FarmCPU, MLM 6.31E-04 1.54E-04 7.070
AX-111802919 4D 508.503,834 FarmCPU, MLM+Q+K 7.58E-04 2.17E-04 15.330
LS-TFW AX-111660453 1A 574.476,865 FarmCPU, MLM, MLM+Q+K 8.01E-04 1.08E-04 6.489
AX-109551739 1A 574.486,675 FarmCPU, MLM, MLM+Q+K 8.01E-04 1.08E-04 6.489
AX-110920832 3B 44.975,458 FarmCPU, MLM 1.62E-04 1.69E-05 6.000
AX-110979247 3B 45.045,393 FarmCPU, MLM 1.77E-04 1.91E-05 6.000
AX-111518365 3B 45.289,556 FarmCPU, MLM 2.02E-04 2.29E-05 5.830
AX-109940356 3B 45.257,506 FarmCPU, MLM 2.27E-04 2.69E-05 5.850
LS-TDW AX-111660453 1A 574.476,865 FarmCPU, MLM, MLM+Q+K 1.09E-04 1.86E-05 8.634
AX-109551739 1A 574.486,675 FarmCPU, MLM, MLM+Q+K 1.09E-04 1.86E-05 8.634
AX-110920832 3B 44.975,458 FarmCPU, MLM 1.95E-04 5.14E-05 6.660
AX-110979247 3B 45.045,393 FarmCPU, MLM 1.97E-04 5.21E-05 6.770
AX-111518365 3B 45.289,556 FarmCPU, MLM 1.90E-04 4.97E-05 7.060
AX-109940356 3B 45.257,506 FarmCPU, MLM 2.98E-04 8.76E-05 6.370
AX-110548993 3B 65.160,500 FarmCPU, MLM 3.61E-04 1.11E-04 7.880
LS-SNA AX-111660453 1A 574.476,865 FarmCPU, MLM, MLM+Q+K 2.93E-04 1.20E-05 7.573
AX-109551739 1A 574.486,675 FarmCPU, MLM, MLM+Q+K 2.93E-04 1.20E-05 7.573
AX-111802919 4D 508.503,834 FarmCPU, MLM+Q+K 1.40E-04 3.17E-05 5.760
LS-TNA AX-111660453 1A 574.476,865 FarmCPU, MLM, MLM+Q+K 2.38E-04 1.92E-05 7.866
AX-109551739 1A 574.486,675 FarmCPU, MLM, MLM+Q+K 2.38E-04 1.92E-05 7.866
AX-111802919 4D 508.503,834 FarmCPU, MLM+Q+K 5.44E-04 3.93E-04 7.030
LS-TNUE AX-110920832 3B 44.975,458 FarmCPU, MLM 8.18E-05 3.36E-05 8.040
AX-110979247 3B 45.045,393 FarmCPU, MLM 2.48E-04 1.26E-04 7.010
LS-TNUE AX-111518365 3B 45.289,556 FarmCPU, MLM 5.61E-05 2.12E-05 8.770
AX-109940356 3B 45.257,506 FarmCPU, MLM 1.36E-04 6.21E-05 7.410
AX-110548993 3B 65.160,500 FarmCPU, MLM 6.67E-04 4.02E-04 9.050
LS-SNUE AX-110920832 3B 44.975,458 FarmCPU, MLM 2.60E-04 1.39E-04 5.980
AX-110979247 3B 45.045,393 FarmCPU, MLM 5.78E-04 3.51E-04 5.320
AX-111518365 3B 45.289,556 FarmCPU, MLM 1.39E-04 6.59E-05 6.910
AX-109940356 3B 45.257,506 FarmCPU, MLM 3.76E-04 2.14E-04 5.570
AX-110548993 3B 65.160,500 FarmCPU, MLM 5.88E-04 3.59E-04 6.670
LS-RFW AX-110920832 3B 44.975,458 FarmCPU, MLM 3.72E-04 6.98E-05 5.660
AX-110979247 3B 45.045,393 FarmCPU, MLM 3.57E-04 6.62E-05 5.770
AX-111518365 3B 45.289,556 FarmCPU, MLM 2.96E-04 5.16E-05 5.690
AX-109940356 3B 45.257,506 FarmCPU, MLM 3.68E-04 6.87E-05 5.840

图5

小麦苗期氮效率性状显著SNP位点的等位基因效应分析 缩写同表1。图a和图b分别表示SNP位点AX-111802919、AX-110548993的等位基因效应分析, 其中A、C、G代表选取相应SNP位点不同基因型。*、**和***分别表示在0.05、0.01和0.001水平差异显著。"

附表2

候选基因及其功能注释"

候选基因
Candidate genes
物理位置
Position (bp)
功能注释
Gene annotations
TraesCS3B02G091100 Chr. 3B: 60250946-60254590 钾刺激焦磷酸盐激活钠泵蛋白
K-stimulated pyrophosphate-energized sodium pump protein
TraesCS3B02G091200 Chr. 3B: 60254745-60262160 脂肪酸氧化复合物亚基α Fatty acid oxidation complex subunit alpha
TraesCS3B02G091300 Chr. 3B: 60271263-60272660 皮层素结合蛋白2 Cortactin-binding protein 2
TraesCS3B02G091400 Chr. 3B: 60365231-60368062 表达的细胞色素P450家族蛋白 Cytochrome P450 family protein, expressed
TraesCS3B02G091500 Chr. 3B: 60548536-60550361 DDT结构域超家族 DDT domain superfamily
TraesCS3B02G091600 Chr. 3B: 60629463-60630222 F-box蛋白 F-box protein
TraesCS3B02G091700 Chr. 3B: 60641814-60642605 可溶性球蛋白 CsAtPR5
TraesCS3B02G091800 Chr. 3B: 60863814-60868553 短链脱氢酶/还原酶家族蛋白 Short-chain dehydrogenase/reductase family protein
TraesCS3B02G091900 Chr. 3B: 61048079-61052327 含XH / XS结构域蛋白 XH/XS domain-containing protein
TraesCS3B02G092000 Chr. 3B: 61098661-61099299 含XH / XS结构域蛋白 XH/XS domain-containing protein
TraesCS3B02G092100 Chr. 3B: 61128394-61128941 DNA拓扑异构酶 DNA topoisomerase
TraesCS3B02G092200 Chr. 3B: 61527695-61528392 SKP1 - like蛋白 SKP1-like protein
TraesCS3B02G092300 Chr. 3B: 61532394-61532792 Na (+)转位NADH-醌还原酶亚基A
Na(+)-translocating NADH-quinone reductase subunit A
TraesCS3B02G092400 Chr. 3B: 61552988-61555381 五肽重复蛋白Pentatricopeptide repeat-containing protein
TraesCS3B02G092500 Chr. 3B: 61560637-61564731 碱性螺旋-环-螺旋(bHLH) DNA结合超家族蛋白
basic helix-loop-helix (bHLH) DNA-binding superfamily protein
TraesCS3B02G092600 Chr. 3B: 61875981-61876268 含B3结构域蛋白 B3 domain-containing protein
TraesCS3B02G092700 Chr. 3B: 61880754-61884071 转移酶家族蛋白 Transferase family protein
TraesCS3B02G092800 Chr. 3B: 61895861-61896952 含NAC结构域的蛋白质 NAC domain-containing protein
TraesCS3B02G092900 Chr. 3B: 61914874-61916355 含NAC结构域的蛋白质 NAC domain-containing protein
TraesCS3B02G093000 Chr. 3B: 62010635-62012186 应激反应蛋白 Stress responsive protein
TraesCS3B02G093100 Chr. 3B: 62214759-62217472 应激反应蛋白 Stress responsive protein
TraesCS3B02G093200 Chr. 3B: 62292476-62293280 周质海藻糖酶 Periplasmic trehalase
TraesCS3B02G093300 Chr. 3B: 62313202-62315456 NAC结构域蛋白 NAC domain protein
TraesCS3B02G093400 Chr. 3B: 62392719-62396533 E3泛素连接酶家族蛋白 E3 Ubiquitin ligase family protein
TraesCS3B02G093500 Chr. 3B: 62415051-62418811 含有BRO1样结构域的内体靶向蛋白
Endosomal targeting BRO1-like domain-containing protein
TraesCS3B02G093600 Chr. 3B: 62530132-62530658 核糖核苷二磷酸还原酶Ribonucleoside-diphosphate reductase
TraesCS3B02G093700 Chr. 3B: 62667563-62672566 丝氨酸蛋白酶HtrA样蛋白 Serine protease HtrA-like
TraesCS3B02G093800 Chr. 3B: 62945115-62948759 类受体激酶 Receptor-like kinase
TraesCS3B02G093900 Chr. 3B: 62948630-62953476 DNA导向的RNA聚合酶β亚基 DNA-directed RNA polymerase subunit beta
TraesCS3B02G094000 Chr. 3B: 62953766-62958847 整合子复合体亚基7 Integrator complex subunit 7
TraesCS3B02G094100 Chr. 3B: 62964339-62972385 受体激酶1 receptor kinase 1
TraesCS3B02G094200 Chr. 3B: 62982063-62983361 包含蛋白的F-box结构域 F-box domain containing protein
TraesCS3B02G094300 Chr. 3B: 62985132-62986025 可溶性球蛋白 CsAtPR5
TraesCS3B02G094400 Chr. 3B: 63107268-63118230 泛素家族蛋白 Ubiquitin family protein
TraesCS3B02G094500 Chr. 3B: 63387316-63390082 F - box蛋白PP2 F-box protein PP2
TraesCS3B02G094700 Chr. 3B: 63392218-63395735 DNA/RNA结合蛋白KIN17 DNA/RNA-binding protein KIN17
TraesCS3B02G094600 Chr. 3B: 63392249-63393457 甾醇-8, 7-异构酶 Sterol-8, 7-isomerase
TraesCS3B02G094800 Chr. 3B: 63398367-63399534 凝集素受体激酶 Lectin receptor kinase
TraesCS3B02G094900 Chr. 3B: 63404102-63406543 假定跨膜蛋白(DUF594) Transmembrane protein, putative (DUF594)
TraesCS3B02G095100 Chr. 3B: 63468035-63472262 单胺氧化酶A阻遏子R1蛋白的锌指结构域
Zinc-finger domain of monoamine-oxidase A repressor R1 protein
TraesCS3B02G095200 Chr. 3B: 63987407-64004822 受体激酶2 Receptor kinase 2
TraesCS3B02G095300 Chr. 3B: 64165574-64178598 受体激酶1 Receptor kinase 1
TraesCS3B02G095400 Chr. 3B: 64242418-64243279 导向蛋白 Dirigent protein
TraesCS3B02G095500 Chr. 3B: 64244480-64249451 氧化还原酶Tas aldo / 酮基还原酶家族蛋白
Oxidoreductase Tas aldo/keto reductase family protein
TraesCS3B02G095700 Chr. 3B: 64298721-64299258 防卫素 Defensin
TraesCS3B02G095800 Chr. 3B: 64306545-64307612 F-box家族蛋白 F-box family protein
TraesCS3B02G096000 Chr. 3B: 64314488-64320468 主要协助转运超家族蛋白 Major facilitator superfamily protein
TraesCS3B02G095900 Chr. 3B: 64314648-64316600 假定受体蛋白激酶 Receptor protein kinase, putative
TraesCS3B02G096100 Chr. 3B: 64413796-64414790 丝氨酸/苏氨酸-蛋白磷酸酶 Serine/threonine-protein phosphatase
TraesCS3B02G096200 Chr. 3B: 64415992-64416598 光调控蛋白 Light-regulated protein
TraesCS3B02G096300 Chr. 3B: 64499581-64500175 重金属相关蛋白 Heavy metal-associated protein
TraesCS3B02G096400 Chr. 3B: 64504291-64508081 带电荷的多泡体蛋白5 Charged multivesicular body protein 5
TraesCS3B02G096500 Chr. 3B: 64508601-64509210 光调控蛋白 Light-regulated protein
TraesCS3B02G096600 Chr. 3B: 64509653-64510875 丝氨酸/苏氨酸-蛋白磷酸酶 Serine/threonine-protein phosphatase
TraesCS3B02G096700 Chr. 3B: 64538075-64539934 多元醇转运子 Polyol transporter
TraesCS3B02G096800 Chr. 3B: 64596330-64597558 丝氨酸/苏氨酸-蛋白磷酸酶 Serine/threonine-protein phosphatase
TraesCS3B02G096900 Chr. 3B: 64695774-64700203 抗病蛋白(NBS-LRR类)家族 Disease resistance protein (NBS-LRR class) family
TraesCS3B02G097000 Chr. 3B: 64700982-64706088 DNA修复蛋白样蛋白 DNA repair protein-like protein
TraesCS3B02G097100 Chr. 3B: 64811084-64813465 E3泛素蛋白连接酶SINA-like 10 E3 ubiquitin-protein ligase SINA-like 10
TraesCS3B02G097200 Chr. 3B: 65193781-65196227 易位子相关蛋白亚基β Translocon-associated protein subunit beta
TraesCS3B02G097300 Chr. 3B: 65198470-65199985 富含半胱氨酸的受体激酶样蛋白 Cysteine-rich receptor-kinase-like protein
TraesCS3B02G097400 Chr. 3B: 65211236-65215663 蛋白质腺毛双折射 Protein trichome birefringence
TraesCS3B02G097500 Chr. 3B: 65268601-65269447 Harpin诱导蛋白 Harpin-induced protein
TraesCS3B02G097600 Chr. 3B: 65335466-65339154 类F-box家族蛋白 F-box family protein-like
TraesCS3B02G097700 Chr. 3B: 65346987-65349183 类PapD超家族蛋白 PapD-like superfamily protein
TraesCS3B02G097800 Chr. 3B: 65358509-65360980 莽草酸激酶家族蛋白 Shikimate kinase family protein
TraesCS3B02G097900 Chr. 3B: 65366570-65369381 缝隙连接β - 4蛋白亚型1 Gap junction beta-4 protein isoform 1
TraesCS3B02G098000 Chr. 3B: 65547132-65548442 叶绿体类囊体腔17.9 kD蛋白 Thylakoid lumenal 17.9 kD protein, chloroplastic
TraesCS3B02G098100 Chr. 3B: 65552302-65554643 非特异性磷脂酶C Non-specific phospholipase C
TraesCS3B02G098200 Chr. 3B: 65558254-65558466 假定跨膜蛋白(DUF 3339) transmembrane protein, putative (DUF 3339)
TraesCS3B02G098300 Chr. 3B: 65560152-65560370 假定跨膜蛋白(DUF 3339) transmembrane protein, putative (DUF 3339)
TraesCS3B02G098400 Chr. 3B: 65560604-65564912 内-贝壳杉烯酸羟化酶2 Ent-kaurenoic acid hydroxylase 2
TraesCS3B02G098500 Chr. 3B: 66050390-66055387 核糖体蛋白L29家族蛋白 Ribosomal protein L29 family protein
TraesCS3B02G098600 Chr. 3B: 66170761-66173676 细胞色素P450家族蛋白 Cytochrome P450 family protein
TraesCS3B02G098700 Chr. 3B: 66180176-66182333 伴侣蛋白dna J Chaperone protein dnaJ
TraesCS3B02G098800 Chr. 3B: 66186166-66190004 RNA识别基序家族蛋白 RNA recognition motif-containing family protein
TraesCS3B02G098900 Chr. 3B: 66416023-66420917 蛋白激酶家族蛋白 Protein kinase family protein
TraesCS3B02G099000 Chr. 3B: 66470014-66471136 木葡聚糖转移/水解酶 Xyloglucan endotransglucosylase/hydrolase
TraesCS3B02G099100 Chr. 3B: 66477677-66478861 木葡聚糖转移/水解酶 Xyloglucan endotransglucosylase/hydrolase
TraesCS3B02G099200 Chr. 3B: 66574594-66577599 核转运蛋白Snurportin-1 Snurportin-1
TraesCS3B02G099300 Chr. 3B: 66577194-66579234 烯醇化酶-磷酸酶E1 Enolase-phosphatase E1
TraesCS3B02G099400 Chr. 3B: 66579716-66581682 蛋白酶HtpX Protease HtpX
TraesCS3B02G099500 Chr. 3B: 66586170-66588662 含五肽重复家族蛋白 Pentatricopeptide repeat-containing family protein
TraesCS3B02G099600 Chr. 3B: 66597468-66603123 细胞色素P450家族蛋白 Cytochrome P450 family protein
TraesCS3B02G099700 Chr. 3B: 66611483-66613539 细胞色素P450家族蛋白 Cytochrome P450 family protein
TraesCS3B02G099800 Chr. 3B: 66678779-66679843 转移酶, 转移糖基 Transferase, transferring glycosyl groups
TraesCS3B02G099900 Chr. 3B: 66706917-66711122 细胞色素P450家族蛋白 Cytochrome P450 family protein
TraesCS3B02G100000 Chr. 3B: 66712696-66714756 细胞色素P450家族蛋白 Cytochrome P450 family protein
TraesCS3B02G100100 Chr. 3B: 66797759-66798919 转移酶, 转移糖基 Transferase, transferring glycosyl groups
TraesCS3B02G100200 Chr. 3B: 66801624-66805418 含五肽重复家族蛋白 Pentatricopeptide repeat-containing family protein
TraesCS3B02G100300 Chr. 3B: 66816983-66820910 FBD相关F-box蛋白 FBD-associated F-box protein
TraesCS3B02G100400 Chr. 3B: 66864005-66871291 阻抑蛋白 Prohibitin
TraesCS3B02G100500 Chr. 3B: 66873290-66877508 NADPH -细胞色素P450还原酶 NADPH-cytochrome P450 reductase
TraesCS3B02G100600 Chr. 3B: 66877543-66882500 谷氨酸脱羧酶 Glutamate decarboxylase
TraesCS3B02G100700 Chr. 3B: 66953395-66958021 假定BZIP转录因子(DUF1664) bZIP transcription factor, putative (DUF1664)
TraesCS3B02G100800 Chr. 3B: 67060640-67062536 30S核糖体蛋白S7 30S ribosomal protein S7
TraesCS3B02G100900 Chr. 3B: 67689687-67690425 含CCHC结构域的锌指蛋白4 Zinc finger CCHC domain-containing protein 4
TraesCS3B02G101000 Chr. 3B: 67944264-67946258 应激反应α-β桶状结构域蛋白 Stress responsive alpha-beta barrel domain protein
TraesCS3B02G101100 Chr. 3B: 68253426-68254192 白细胞介素-4 Interleukin-4
TraesCS3B02G101200 Chr. 3B: 68348282-68356139 单铜氧化酶样蛋白SKU5 Monocopper oxidase-like protein SKU5
TraesCS3B02G101300 Chr. 3B: 68398778-68406173 倍半萜合酶 Sesquiterpene synthase
TraesCS3B02G101400 Chr. 3B: 68427410-68429508 细胞色素P450 Cytochrome P450
TraesCS3B02G101500 Chr. 3B: 68442260-68445404 单铜氧化酶样蛋白SKU5 Monocopper oxidase-like protein SKU5
TraesCS3B02G101600 Chr. 3B: 68450231-68453892 跨膜蛋白 Transmembrane protein
TraesCS3B02G101700 Chr. 3B: 68463718-68468382 CC-NBS-LRR家族抗病蛋白 CC-NBS-LRR family disease resistance protein
TraesCS3B02G101900 Chr. 3B: 68588515-68591362 R3H结构域含有蛋白 R3H domain containing protein
TraesCS3B02G102000 Chr. 3B: 68593361-68593936 CDNA, 克隆号: J090004K22, 全插入序列
cDNA, clone: J090004K22, full insert sequence
TraesCS3B02G102100 Chr. 3B: 68871684-68880041 脂肪酸氧化复合物亚基α Fatty acid oxidation complex subunit alpha
TraesCS3B02G102200 Chr. 3B: 68881218-68885390 蛋白AATF Protein AATF
TraesCS3B02G102300 Chr. 3B: 68899038-68902413 剪接因子 Splicing factor
TraesCS3B02G102400 Chr. 3B: 68903863-68910705 TBC1结构域家族成员 TBC1 domain family member
TraesCS3B02G102500 Chr. 3B: 68910939-68914684 半胱氨酸合成酶 Cysteine synthase
TraesCS3B02G102600 Chr. 3B: 69351976-69360003 NADP依赖的α,β-不饱和醛酮双键还原酶
NADP-dependent alkenal double bond reductase
TraesCS3B02G102700 Chr. 3B: 69361029-69362107 F-box家族蛋白 F-box family protein
TraesCS3B02G102800 Chr. 3B: 69603488-69606079 嘧啶核苷特异性核糖核苷水解酶 Pyrimidine-specific ribonucleoside hydrolase
TraesCS3B02G102900 Chr. 3B: 69605986-69609368 类AT2G31890蛋白 AT2G31890-like protein
TraesCS3B02G103000 Chr. 3B: 69620569-69621345 F-box家族蛋白 F-box family protein
TraesCS3B02G103100 Chr. 3B: 69643792-69645390 GDSL酯酶/脂肪酶 GDSL esterase/lipase
TraesCS3B02G103200 Chr. 3B: 69811561-69818149 转录因子GTE9 Transcription factor GTE9
TraesCS3B02G103300 Chr. 3B: 70092982-70093620 RING/U-box超家族蛋白 RING/U-box superfamily protein
TraesCS4D02G351800 Chr. 4D: 503505981-503507640 短链脱氢酶/还原酶家族蛋白 Short-chain dehydrogenase/reductase family protein
TraesCS4D02G351900 Chr. 4D: 503563204-503565726 假定细胞色素P450 Cytochrome P450, putative
TraesCS4D02G352000 Chr. 4D: 503596853-503599004 冷调节蛋白27 Cold regulated protein 27
TraesCS4D02G352100 Chr. 4D: 503606966-503607924 假定FAD结合小檗碱家族蛋白 FAD-binding Berberine family protein, putative
TraesCS4D02G352200 Chr. 4D: 503615478-503621888 假定呼吸爆发氧化酶 Respiratory burst oxidase, putative
TraesCS4D02G352300 Chr. 4D: 503622240-503624321 F-box家族蛋白 F-box family protein
TraesCS4D02G352400 Chr. 4D: 503668321-503670497 类CASP蛋白 CASP-like protein
TraesCS4D02G352500 Chr. 4D: 503672009-503675192 假定受体激酶 Receptor-kinase, putative
TraesCS4D02G352600 Chr. 4D: 503742338-503747796 锚蛋白重复蛋白家族样蛋白 Ankyrin repeat protein family-like protein
TraesCS4D02G352700 Chr. 4D: 503785960-503789641 类受体激酶 Receptor kinase-like protein
TraesCS4D02G352800 Chr. 4D: 503791201-503793287 吡哆醇/吡哆胺5'-磷酸氧化酶 Pyridoxine/pyridoxamine 5'-phosphate oxidase
TraesCS4D02G352900 Chr. 4D: 503804138-503809364 假定WD-重复蛋白 WD-repeat protein, putative
TraesCS4D02G353000 Chr. 4D: 503905849-503906196 环核苷酸门控通道9 Cyclic nucleotide gated channel 9
TraesCS4D02G353100 Chr. 4D: 503918550-503920992 L-古洛糖酸内酯氧化酶 L-gulonolactone oxidase
TraesCS4D02G353200 Chr. 4D: 503964657-503967442 五肽重复蛋白 Pentatricopeptide repeat-containing protein
TraesCS4D02G353300 Chr. 4D: 503968880-503969956 BTB / POZ和MATH结构域蛋白2 BTB/POZ and MATH domain-containing protein 2
TraesCS4D02G353400 Chr. 4D: 503997489-503999419 类细胞色素P450蛋白 Cytochrome P450-like protein
TraesCS4D02G353500 Chr. 4D: 504043183-504046044 L-古洛糖酸内酯氧化酶 L-gulonolactone oxidase
TraesCS4D02G353600 Chr. 4D: 504073956-504079603 双功能岩藻糖激酶/岩藻糖焦磷酸化酶
Bifunctional fucokinase/fucose pyrophosphorylase
TraesCS4D02G353700 Chr. 4D: 504079683-504081584 吡哆醇/吡哆胺5'-磷酸氧化酶 Pyridoxine/pyridoxamine 5'-phosphate oxidase
TraesCS4D02G353800 Chr. 4D: 504087839-504093419 双特异性磷酸酶 Dual specificity phosphatase
TraesCS4D02G353900 Chr. 4D: 504097765-504100105 细胞周期蛋白依赖性激酶抑制剂SMR1-like蛋白
Cyclin-dependent kinase inhibitor SMR1-like protein
TraesCS4D02G354000 Chr. 4D: 504108998-504111321 UPF0548蛋白 UPF0548 protein
TraesCS4D02G354100 Chr. 4D: 504114728-504115931 含有LOB结构域的蛋白 LOB domain-containing protein
TraesCS4D02G354200 Chr. 4D: 504177198-504178305 含有LOB结构域的蛋白 LOB domain-containing protein
TraesCS4D02G354300 Chr. 4D: 504197596-504200977 RING / U - box超家族蛋白 RING/U-box superfamily protein
TraesCS4D02G354400 Chr. 4D: 504261950-504265251 NBS-LRR抗病蛋白 NBS-LRR disease resistance protein
TraesCS4D02G354500 Chr. 4D: 504266625-504267882 BTB/POZ结构域含有蛋白 BTB/POZ domain containing protein
TraesCS4D02G354600 Chr. 4D: 504271326-504271613 B3结构域蛋白家族 B3 domain-containing protein family
TraesCS4D02G354700 Chr. 4D: 504273665-504281645 含有蛋白的F - box和Leucine Rich Repeat结构域, 推测为异构体1
F-box and Leucine Rich Repeat domains containing protein, putative isoform 1
TraesCS4D02G354800 Chr. 4D: 504315254-504317909 小核核糖核蛋白样蛋白 Small nuclear ribonucleoprotein-like protein
TraesCS4D02G354900 Chr. 4D: 504328740-504331280 硅转运蛋白 Silicon transporter
TraesCS4D02G355000 Chr. 4D: 504339499-504344372 SKP1家族蛋白 SKP1 family protein
TraesCS4D02G355100 Chr. 4D: 504344908-504345720 扩张素蛋白 Expansin protein
TraesCS4D02G355200 Chr. 4D: 504508069-504508389 开花促进因子1样蛋白1 Flowering-promoting factor 1-like protein 1
TraesCS4D02G355300 Chr. 4D: 504511651-504512007 花粉类过敏原 Pollen allergen-like protein
TraesCS4D02G355400 Chr. 4D: 504521809-504522099 丝氨酸/苏氨酸-蛋白激酶 Serine/threonine-protein kinase
TraesCS4D02G355500 Chr. 4D: 504562124-504562480 花粉类过敏原 Pollen allergen-like protein
TraesCS4D02G355600 Chr. 4D: 504597403-504598281 水孔蛋白 Aquaporin
TraesCS4D02G355700 Chr. 4D: 504996446-504997582 扩张素蛋白 Expansin protein
TraesCS4D02G355800 Chr. 4D: 505188396-505189587 扩张素蛋白 Expansin protein
TraesCS4D02G355900 Chr. 4D: 505192205-505197656 类SKP1蛋白21 SKP1-like protein 21
TraesCS4D02G356000 Chr. 4D: 505237477-505237851 花粉类过敏原 Pollen allergen-like protein
TraesCS4D02G356100 Chr. 4D: 505307336-505307578 液泡分选受体2 VACUOLAR SORTING RECEPTOR 2
TraesCS4D02G356200 Chr. 4D: 505325871-505329827 支链氨基酸氨基转移酶 Branched-chain-amino-acid aminotransferase
TraesCS4D02G356300 Chr. 4D: 505330644-505333517 脂肪酰辅酶A还原酶 Fatty acyl-CoA reductase
TraesCS4D02G356400 Chr. 4D: 505352502-505354843 E3泛素连接酶 E3 ubiquitin-protein ligase
TraesCS4D02G356500 Chr. 4D: 505357085-505359092 F-box / RNI-like超家族蛋白 F-box/RNI-like superfamily protein
TraesCS4D02G356600 Chr. 4D: 505363557-505367310 烷基转移酶 Alkyl transferase
TraesCS4D02G356700 Chr. 4D: 505381091-505384995 F-box家族蛋白 F-box family protein
TraesCS4D02G356800 Chr. 4D: 505388341-505393599 假定表达的PAIR1蛋白 PAIR1 protein, putative, expressed
TraesCS4D02G356900 Chr. 4D: 505427738-505432457 磷脂酰肌醇N -乙酰氨基葡萄糖转移酶亚基P样蛋白
Phosphatidylinositol N-acetyglucosaminlytransferase subunit P-like protein
TraesCS4D02G357000 Chr. 4D: 505432671-505433054 碱性螺旋-环-螺旋( b HLH ) DNA结合超家族蛋白
Basic helix-loop-helix (bHLH) DNA-binding superfamily protein
TraesCS4D02G357100 Chr. 4D: 505433675-505437130 腺苷三磷酸双磷酸酶 Apyrase
TraesCS4D02G357200 Chr. 4D: 505498001-505500622 酸性磷酸酶1 Acid phosphatase 1
TraesCS4D02G357300 Chr. 4D: 505514818-505517173 酸性磷酸酶1 Acid phosphatase 1
TraesCS4D02G357400 Chr. 4D: 505553303-505554841 酸性磷酸酶1 Acid phosphatase 1
TraesCS4D02G357500 Chr. 4D: 505700443-505700892 抗病蛋白( TIR - NBS - LRR类)家族
Disease resistance protein (TIR-NBS-LRR class) family
TraesCS4D02G357600 Chr. 4D: 505725280-505725876 代谢型谷氨酸受体2 Metabotropic glutamate receptor 2
TraesCS4D02G357700 Chr. 4D: 505983762-505988144 RING / U - box超家族蛋白 RING/U-box superfamily protein
TraesCS4D02G357800 Chr. 4D: 505988576-505992394 假定蛋白磷酸酶2c Protein phosphatase 2c, putative
TraesCS4D02G357900 Chr. 4D: 505998197-506003740 BAG家族分子伴侣调节因子6 BAG family molecular chaperone regulator 6
TraesCS4D02G358000 Chr. 4D: 506029864-506030373 导向蛋白 Dirigent protein
TraesCS4D02G358100 Chr. 4D: 506050808-506053015 间天冬氨酸酶-1 Metacaspase-1
TraesCS4D02G358200 Chr. 4D: 506061720-506062465 环指蛋白44 RING finger protein 44
TraesCS4D02G358300 Chr. 4D: 506064397-506070542 抗病蛋白RPM1 Disease resistance protein RPM1
TraesCS4D02G358400 Chr. 4D: 506072327-506074686 抗病蛋白RPM1 Disease resistance protein RPM1
TraesCS4D02G358500 Chr. 4D: 506103285-506105538 多酚氧化酶 Polyphenol oxidase
TraesCS4D02G358600 Chr. 4D: 506130446-506133842 液泡蛋白分选相关蛋白2 1 Vacuolar protein sorting-associated protein 2 1
TraesCS4D02G358700 Chr. 4D: 506204779-506206379 木葡聚糖转移/水解酶 Xyloglucan endotransglucosylase/hydrolase
TraesCS4D02G358800 Chr. 4D: 506285006-506287456 间天冬氨酸酶-I Metacaspase-1
TraesCS4D02G358900 Chr. 4D: 506339104-506340233 热激转录因子 Heat shock transcription factor
TraesCS4D02G359000 Chr. 4D: 506463560-506466966 ATP依赖的RNA解旋酶 ATP dependent RNA helicase
TraesCS4D02G359100 Chr. 4D: 506605746-506607107 MYB转录因子 MYB transcription factor
TraesCS4D02G359200 Chr. 4D: 506720096-506722226 多酚氧化酶 Polyphenol oxidase
TraesCS4D02G359300 Chr. 4D: 506746643-506747502 导向蛋白 Dirigent protein
TraesCS4D02G359400 Chr. 4D: 506852817-506854686 酪氨酸脱羧酶 Tyrosine decarboxylase
TraesCS4D02G359500 Chr. 4D: 506871482-506876195 糖基转移酶 Glycosyltransferase
TraesCS4D02G359600 Chr. 4D: 506967357-506975612 同源盒亮氨酸拉链蛋白 Homeobox leucine-zipper protein
TraesCS4D02G359700 Chr. 4D: 507001378-507002738 酪氨酸脱羧酶 Tyrosine decarboxylase
TraesCS4D02G359800 Chr. 4D: 507005481-507007335 胍基丁胺香豆酰基转移酶-2 Agmatine coumaroyltransferase-2
TraesCS4D02G359900 Chr. 4D: 507119357-507119950 14 k Da富含脯氨酸蛋白DC2.15 14 kDa proline-rich protein DC2.15
TraesCS4D02G360000 Chr. 4D: 507124730-507125155 14 k Da富含脯氨酸蛋白DC2.15 14 kDa proline-rich protein DC2.15
TraesCS4D02G360100 Chr. 4D: 507163625-507164068 14 k Da富含脯氨酸蛋白DC2.15 14 kDa proline-rich protein DC2.15
TraesCS4D02G360200 Chr. 4D: 507271557-507274006 稻纵卷叶螟转录因子家族蛋白 Scarecrow transcription factor family protein
TraesCS4D02G360300 Chr. 4D: 507279851-507284820 信号识别颗粒蛋白 Signal recognition particle protein
TraesCS4D02G360400 Chr. 4D: 507348199-507352544 F-box蛋白 F-box protein
TraesCS4D02G360500 Chr. 4D: 507417297-507423921 抗病蛋白RPM1 Disease resistance protein RPM1
TraesCS4D02G360600 Chr. 4D: 507559405-507560809 MYB相关转录因子 MYB-related transcription factor
TraesCS4D02G360700 Chr. 4D: 507665144-507666913 HXXXD型酰基转移酶家族蛋白 HXXXD-type acyl-transferase family protein
TraesCS4D02G360800 Chr. 4D: 507679722-507683815 黄素依赖型单加氧酶 Flavin-containing monooxygenase
TraesCS4D02G360900 Chr. 4D: 507686127-507689164 黄素依赖型单加氧酶 Flavin-containing monooxygenase
TraesCS4D02G361000 Chr. 4D: 507712341-507717027 黄素依赖型单加氧酶 Flavin-containing monooxygenase
TraesCS4D02G361100 Chr. 4D: 507890203-507895188 假定T-box转录因子(DUF863) T-box transcription factor, putative (DUF863)
TraesCS4D02G361200 Chr. 4D: 507901581-507902876 类C2H2锌指蛋白 C2H2-like zinc finger protein
TraesCS4D02G361300 Chr. 4D: 507965542-507967822 钠离子转运蛋白家族蛋白 Sodium transporter family protein
TraesCS4D02G361400 Chr. 4D: 508256672-508263207 氧化甾醇结合蛋白 Oxysterol-binding protein
TraesCS4D02G361500 Chr. 4D: 508264326-508271976 硝酸盐转运蛋白1.1 Nitrate transporter 1.1
TraesCS4D02G361600 Chr. 4D: 508339521-508343262 丝氨酸/苏氨酸-蛋白激酶 Serine/threonine-protein kinase
TraesCS4D02G361700 Chr. 4D: 508431333-508432960 扩张素蛋白 Expansin protein
TraesCS4D02G361800 Chr. 4D: 508479234-508480543 非特异性磷脂酶C3 non-specific phospholipase C3
TraesCS4D02G361900 Chr. 4D: 508500284-508502749 扩张素蛋白 Expansin protein
TraesCS4D02G362000 Chr. 4D: 508528075-508528350 推定RING / U - box超家族蛋白 RING/U-box superfamily protein, putative
TraesCS4D02G362100 Chr. 4D: 508529210-508529773 锌指蛋白康斯坦斯-LIKE 1 Zinc finger protein CONSTANS-LIKE 1
TraesCS4D02G362200 Chr. 4D: 508591122-508591336 叶绿体30S核糖体蛋白 30S ribosomal protein S19, chloroplastic
TraesCS4D02G362300 Chr. 4D: 508672464-508681236 2-磷酸甘油酸激酶, 推定, 表达 2-phosphoglycerate kinase, putative, expressed
TraesCS4D02G362400 Chr. 4D: 508682913-508702166 CCR4 - NOT转录复合体亚基1 CCR4-NOT transcription complex subunit 1
TraesCS4D02G362500 Chr. 4D: 508806906-508807847 咖啡酰辅酶A-O-甲基转移酶 Caffeoyl-CoA O-methyltransferase
TraesCS4D02G362600 Chr. 4D: 508809651-508819038 ATP依赖的RNA解旋酶 ATP-dependent RNA helicase
TraesCS4D02G362700 Chr. 4D: 508938139-508938936 伴侣蛋白Clp B Chaperone protein ClpB
TraesCS4D02G362800 Chr. 4D: 509056631-509058516 GATA转录因子 GATA transcription factor
TraesCS4D02G362900 Chr. 4D: 509095354-509096715 基质金属蛋白酶 Matrix metalloproteinase
TraesCS4D02G363000 Chr. 4D: 509098244-509100075 2 -酮戊二酸( 2OG )和Fe ( II )依赖的加氧酶超家族蛋白
2-oxoglutarate (2OG) and Fe(II)-dependent oxygenase superfamily protein
TraesCS4D02G363100 Chr. 4D: 509100136-509101897 2 -酮戊二酸( 2OG )和Fe ( II )依赖的加氧酶超家族蛋白
2-oxoglutarate (2OG) and Fe(II)-dependent oxygenase superfamily protein
TraesCS4D02G363200 Chr. 4D: 509101901-509103877 2 -酮戊二酸( 2OG )和Fe ( II )依赖的加氧酶超家族蛋白
2-oxoglutarate (2OG) and Fe(II)-dependent oxygenase superfamily protein
TraesCS4D02G363300 Chr. 4D: 509104468-509106248 2 -酮戊二酸( 2OG )和Fe ( II )依赖的加氧酶超家族蛋白
2-oxoglutarate (2OG) and Fe(II)-dependent oxygenase superfamily protein
TraesCS4D02G363400 Chr. 4D: 509114738-509121487 PHD锌指蛋白 PHD-finger protein
TraesCS4D02G363500 Chr. 4D: 509134936-509137014 细胞色素P450家族蛋白, 表达 Cytochrome P450 family protein, expressed
TraesCS4D02G363600 Chr. 4D: 509172859-509175128 生长素激活蛋白(DUF828) Auxin canalization protein (DUF828)
TraesCS4D02G363700 Chr. 4D: 509175058-509177724 甲硫氨酰基-t RNA甲酰基转移酶 Methionyl-tRNA formyltransferase
TraesCS4D02G363800 Chr. 4D: 509181987-509182706 假定跨膜蛋白 Transmembrane protein, putative
TraesCS4D02G363900 Chr. 4D: 509184569-509189902 外被体亚基α Coatomer subunit alpha
TraesCS4D02G364000 Chr. 4D: 509224814-509228514 磷脂酰丝氨酸脱羧酶原酶 Phosphatidylserine decarboxylase proenzyme
TraesCS4D02G364100 Chr. 4D: 509236273-509239739 细胞数调节因子6 Cell number regulator 6
TraesCS4D02G364200 Chr. 4D: 509272555-509273889 F - box家族蛋白 F-box family protein
TraesCS4D02G364300 Chr. 4D: 509275914-509277096 F - box家族蛋白 F-box family protein
TraesCS4D02G364400 Chr. 4D: 509282253-509284348 锌指结构蛋白 Zinc finger constans-like protein
TraesCS4D02G364500 Chr. 4D: 509338852-509340992 锌指结构蛋白 Zinc finger constans-like protein
TraesCS4D02G364600 Chr. 4D: 509363547-509372502 推定解旋酶 Helicase, putative
TraesCS4D02G364700 Chr. 4D: 509427073-509429074 细胞色素P450家族蛋白 Cytochrome P450 family protein
TraesCS4D02G364800 Chr. 4D: 509430091-509437414 脂酶/酯酶/硫酯酶家族蛋白 Esterase/lipase/thioesterase family protein
TraesCS4D02G364900 Chr. 4D: 509451626-509454010 细胞核样天冬氨酸蛋白酶 Nucellin-like aspartic protease
TraesCS4D02G365000 Chr. 4D: 509456698-509458706 天冬氨酸蛋白酶Asp1 Aspartic proteinase Asp1
TraesCS4D02G365100 Chr. 4D: 509477168-509481540 鞘磷脂磷酸二酯酶4 Sphingomyelin phosphodiesterase 4
TraesCS4D02G365200 Chr. 4D: 509504323-509504880 蛋白酶体组装伴侣4 Proteasome assembly chaperone 4
TraesCS4D02G365300 Chr. 4D: 509510624-509512063 推定富含亮氨酸重复类受体蛋白激酶家族蛋白
Leucine-rich repeat receptor-like protein kinase family protein, putative
TraesCS4D02G365400 Chr. 4D: 509512085-509518918 富含亮氨酸重复类受体蛋白激酶家族蛋白
Leucine-rich repeat receptor-like protein kinase family protein
TraesCS4D02G365500 Chr. 4D: 509534509-509537103 富含亮氨酸重复类受体蛋白激酶家族蛋白
Leucine-rich repeat receptor-like protein kinase family
TraesCS4D02G365600 Chr. 4D: 509645777-509648952 醇脱氢酶 Alcohol dehydrogenase
TraesCS4D02G365700 Chr. 4D: 509649943-509652123 磷酸甘油酸变位酶家族蛋白 Phosphoglycerate mutase family protein
TraesCS4D02G365800 Chr. 4D: 509662882-509667974 推定糖转运蛋白 Sugar transporter, putative
TraesCS4D02G365900 Chr. 4D: 509669400-509677298 蛋白激酶 Protein kinase
TraesCS4D02G366000 Chr. 4D: 509819115-509827442 蛋白Arabidillo 1 Protein ARABIDILLO 1
TraesCS4D02G366100 Chr. 4D: 509827788-509831298 棕榈酰蛋白硫酯酶包含蛋白, 表达
Palmitoyl protein thioesterase containing protein, expressed
TraesCS4D02G366200 Chr. 4D: 509837611-509839060 糖基转移酶 Glycosyltransferase
TraesCS4D02G366300 Chr. 4D: 509850878-509852835 NBS - LRR抗病蛋白样蛋白 NBS-LRR disease resistance protein-like protein
TraesCS4D02G366400 Chr. 4D: 509852857-509854941 抗病蛋白( NBS - LRR类)家族 Disease resistance protein (NBS-LRR class) family

图6

氮效率相关性状候选基因的表达热量图 红色代表该基因在某组织中表达量高, 蓝色则代表在某组织表达量低。"

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