[1] Chaves M M, Maroco J P, Pereira J S. Understanding plant responses to drought-from genes to the whole plant. Funct Plant Biol, 2003, 30: 239–264.

[2] Todaka D, Zhao Y, Yoshida T, Kudo M, Kidokoro S, Mizoi J, Kodaira K S, Takebayashi Y, Kojima M, Sakakibara H, et al. Temporal and spatial changes in gene expression, metabolite accumulation and phytohormone content in rice seedlings grown under drought stress conditions. Plant J, 2017, 90: 61–78.

[3] 任洪雷, 朱筱, 张丰屹, 张必弦, 王家军, 王金生, 吴俊江, 王广金, 邱丽娟. 干旱胁迫的影响及抗旱性研究进展. 分子植物育种, 网络首发[2024-01-19], http://kns.cnki.net/kcms/detail/46.1068.S.20240119.1548.002.
Ren H L, Zhu X, Zhang F Y, Zhang B X, Wang J J, Wang J S, Wu J J, Wang G J, Qiu L J. Effect of drought stress and research progress of drought resistance. Mol Plant Breed, Published online [2024-01-19], http://kns.cnki.net/kcms/detail/46.1068.S.20240119.1548.002.

[4] 万书波, 封海胜, 王秀贞. 花生营养成分综合评价与产业化发展战略研究. 花生学报, 2004, 33(2): 1–6.
Wan S B, Feng H S, Wang X Z. Synthetically evaluation on peanut nutrients and study on its industrialization development strategy. J Peanut Sci, 2004, 33(2): 1–6 (in Chinese with English abstract).

[5] 孙海艳, 侯乾, 董文召, 王晶珊, 刘立峰, 雷永. 我国花生品种发展现状. 中国种业, 2022, (7): 15–17.
Sun H Y, Hou Q, Dong W Z, Wang J S, Liu L F, Lei Y. Development status of peanut varieties in China. China Seed Ind, 2022, (7): 15–17 (in Chinese with English abstract).

[6] Ding L, Lu Z F, Gao L M, Guo S W, Shen Q R. Is nitrogen a key determinant of water transport and photosynthesis in higher plants upon drought stress? Front Plant Sci, 2018, 9: 1143.

[7] 顾学花, 孙莲强, 高波, 孙奇泽, 刘辰, 张佳蕾, 李向东. 施钙对干旱胁迫下花生生理特性、产量和品质的影响. 应用生态学报, 2015, 26: 1433–1439.
Gu X H, Sun L Q, Gao B, Sun Q Z, Liu C, Zhang J L, Li X D. Effects of calcium fertilizer application on peanut growth, physiological characteristics, yield and quality under drought stress. Chin J Appl Ecol, 2015, 26: 1433–1439 (in Chinese with English abstract).

[8] 武志刚, 武舒佳, 王迎春, 郑琳琳. 植物中钙依赖蛋白激酶(CDPK)的研究进展. 草业学报, 2018, 27(1): 204–214.
Wu Z G, Wu S J, Wang Y C, Zheng L L. Advances in studies of calcium-dependent protein kinase (CDPK) in plants. Acta Pratac Sin, 2018, 27(1): 204–214 (in Chinese with English abstract).

[9] Kudla J, Batistic O, Hashimoto K. Calcium signals: the lead currency of plant information processing. Plant Cell, 2010, 22: 541–563.

[10] Yip Delormel T, Boudsocq M. Properties and functions of calcium-dependent protein kinases and their relatives in Arabidopsis thaliana. New Phytol, 2019, 224: 585–604.

[11] Zhu S Y, Yu X C, Wang X J, Zhao R, Li Y, Fan R C, Shang Y, Du S Y, Wang X F, Wu F Q, et al. Two calcium-dependent protein kinases, CPK4 and CPK11, regulate abscisic acid signal transduction in Arabidopsis. Plant Cell, 2007, 19: 3019–3036.

[12] Bin L H, Xu Z L, Chu Y Q, Yan Y, Nie X J, Song W N. Genome-wide analysis of calcium-dependent protein kinase (CDPK) family and functional characterization of TaCDPK25-U in response to drought stress in wheat. Environ Exp Bot, 2023, 209: 105277.

[13] Yu T F, Zhao W Y, Fu J D, Liu Y W, Chen M, Zhou Y B, Ma Y Z, Xu Z S, Xi Y J. Genome-wide analysis of CDPK family in foxtail millet and determination of SiCDPK24 functions in drought stress. Front Plant Sci, 2018, 9: 651.

[14] Li X D, Gao Y Q, Wu W H, Chen L M, Wang Y. Two calcium-dependent protein kinases enhance maize drought tolerance by activating anion channel ZmSLAC1 in guard cells. Plant Biotechnol J, 2022, 20: 143–157.

[15] Sirover M A. On the functional diversity of glyceraldehyde-3-phosphate dehydrogenase: biochemical mechanisms and regulatory control. Biochim Biophys Acta, 2011, 1810: 741–751.

[16] Rius S P, Casati P, Iglesias A A, Gomez-Casati D F. Characterization of Arabidopsis lines deficient in GAPC-1, a cytosolic NAD-dependent glyceraldehyde-3-phosphate dehydrogenase. Plant Physiol, 2008, 148: 1655–1667.

[17] Anoman A D, Muñoz-Bertomeu J, Rosa-Téllez S, Flores-Tornero M, Serrano R, Bueso E, Fernie A R, Segura J, Ros R. Plastidial glycolytic glyceraldehyde-3-phosphate dehydrogenase is an important determinant in the carbon and nitrogen metabolism of heterotrophic cells in Arabidopsis. Plant Physiol, 2015, 169: 1619–1637.

[18] 王芳, 陈巧红, 董乐, 王云, 朱国立, 许珊珊, 黄苹苹, 王建颖, 林娟. 蓖麻3-磷酸甘油醛脱氢酶的基因克隆及表达分析. 分子植物育种, 2018, 16: 7965–7974.
Wang F, Chen Q H, Dong L, Wang Y, Zhu G L, Xu S S, Huang P P, Wang J Y, Lin J. Cloning and expression analysis of glyceraldehyde-3-phosphate dehydrogenase from castor (Ricinus communis L.). Mol Plant Breed, 2018, 16: 7965–7974 (in Chinese with English abstract).

[19] Simkin A J, Alqurashi M, Lopez-Calcagno P E, Headland L R, Raines C A. Glyceraldehyde-3-phosphate dehydrogenase subunits A and B are essential to maintain photosynthetic efficiency. Plant Physiol, 2023, 192: 2989–3000.

[20] Guo L, Devaiah S P, Narasimhan R, Pan X Q, Zhang Y Y, Zhang W H, Wang X M. Cytosolic glyceraldehyde-3-phosphate dehydrogenases interact with phospholipase Dδ to transduce hydrogen peroxide signals in the Arabidopsis response to stress. Plant Cell, 2012, 24: 2200–2212.

[21] Zeng L F, Deng R, Guo Z P, Yang S S, Deng X P. Genome-wide identification and characterization of Glyceraldehyde-3-phosphate dehydrogenase genes family in wheat (Triticum aestivum). BMC Genomics, 2016, 17: 240.

[22] Vescovi M, Zaffagnini M, Festa M, Trost P, Lo Schiavo F, Costa A. Nuclear accumulation of cytosolic glyceraldehyde-3-phosphate dehydrogenase in cadmium-stressed Arabidopsis roots. Plant Physiol, 2013, 162: 333–346.

[23] Zhao X C, Wang J, Xia N, Qu Y W, Zhan Y H, Teng W L, Li H Y, Li W B, Li Y G, Zhao X, et al. Genome-wide identification and analysis of glyceraldehyde-3-phosphate dehydrogenase family reveals the role of GmGAPDH14 to improve salt tolerance in soybean (Glycine max L.). Front Plant Sci, 2023, 14: 1193044.

[24] 石运庆, 王建, 苗华荣, 胡晓辉, 陈静. 不同花生品种芽期抗旱性筛选及其田间验证. 山东农业科学, 2015, 47(2): 34–37.
Shi Y Q, Wang J, Miao H R, Hu X H, Chen J. Drought resistance screening of different peanut varieties at germination stage and its field validation. Shandong Agric Sci, 2015, 47(2): 34–37 (in Chinese with English abstract).

[25] 王彩, 万勇善, 刘风珍, 张昆. 苗期PEG渗透胁迫条件下花生品种抗旱性的研究. 山东农业科学, 2018, 50(6): 65–71.
Wang C, Wan Y S, Liu F Z, Zhang K. Study on drought resistance of peanut varieties at seedling stage under PEG6000 osmotic stress. Shandong Agric Sci, 2018, 50(6): 65–71 (in Chinese with English abstract).

[26] Jiang C J, Li X L, Zou J X, Ren J Y, Jin C Y, Zhang H, Yu H Q, Jin H. Comparative transcriptome analysis of genes involved in the drought stress response of two peanut (Arachis hypogaea L.) varieties. BMC Plant Biol, 2021, 21: 64.

[27] Li H, Handsaker B, Wysoker A, Fennell T, Ruan J, Homer N, Marth G, Abecasis G, Durbin R, 1000 Genome Project Data Processing Subgroup. The sequence alignment/map format and SAMtools. Bioinformatics, 2009, 25: 2078–2079.

[28] Robinson M D, McCarthy D J, Smyth G K. edgeR: a Bioconductor package for differential expression analysis of digital gene expression data. Bioinformatics, 2010, 26: 139–140.

[29] Chen S F, Zhou Y Q, Chen Y R, Gu J. Fastp: an ultra-fast all-in-one FASTQ preprocessor. Bioinformatics, 2018, 34: i884–i890.

[30] Kim D, Paggi J M, Park C, Bennett C, Salzberg S L. Graph-based genome alignment and genotyping with HISAT2 and HISAT-genotype. Nat Biotechnol, 2019, 37: 907–915.

[31] Chen C J, Chen H, Zhang Y, Thomas H R, Frank M H, He Y H, Xia R. TBtools: an integrative toolkit developed for interactive analyses of big biological data. Mol Plant, 2020, 13: 1194–1202.

[32] Yang S, Wang J G, Tang Z H, Li Y, Zhang J L, Guo F, Meng J J, Cui F, Li X G, Wan S B. Calcium/calmodulin modulates salt responses by binding a novel interacting protein SAMS1 in peanut (Arachis hypogaea L.). Crop J, 2023, 11: 21–32.

[33] Livak K J, Schmittgen T D. Analysis of relative gene expression data using real-time quantitative PCR and the 2^−ΔΔCT method. Methods, 2001, 25: 402–408.

[34] Potter S C, Luciani A, Eddy S R, Park Y, Lopez R, Finn R D. HMMER web server: 2018 update. Nucleic Acids Res, 2018, 46: W200–W204.

[35] Bailey T L, Johnson J, Grant C E, Noble W S. The MEME suite. Nucleic Acids Res, 2015, 43: W39–W49.

[36] Spitz F, Furlong E E M. Transcription factors: from enhancer binding to developmental control. Nat Rev Genet, 2012, 13: 613–626.

[37] Kumar S, Stecher G, Tamura K. MEGA7: molecular evolutionary genetics analysis version 7.0 for bigger datasets. Mol Biol Evol, 2016, 33: 1870–1874.

[38] Letunic I, Bork P. Interactive tree of life (iTOL) v5: an online tool for phylogenetic tree display and annotation. Nucleic Acids Res, 2021, 49: W293–W296.

[39] Deng Y H, Chen Q J, Qu Y Y. Protein S-acyl transferase GhPAT27 was associated with Verticillium wilt resistance in cotton. Plants, 2022, 11: 2758.

[40] Li Q Z, Hu T, Lu T J, Yu B, Zhao Y. Calcium-dependent protein kinases CPK3/4/6/11 and 27 respond to osmotic stress and activate SnRK2s in Arabidopsis. Dev Cell, 2025, 60: 1423–1438.e8.

[41] 刘小龙, 李霞, 钱宝云, 唐玉婷. 植物体内钙信号及其在调节干旱胁迫中的作用. 西北植物学报, 2014, 34: 1927–1936.
Liu X L, Li X, Qian B Y, Tang Y T. Ca²⁺ signal transduction and its regulation role under drought stress in plant. Acta Bot Boreali-Occident Sin, 2014, 34: 1927–1936 (in Chinese with English abstract).

[42] Morigasaki S, Shimada K, Ikner A, Yanagida M, Shiozaki K. Glycolytic enzyme GAPDH promotes peroxide stress signaling through multistep phosphorelay to a MAPK cascade. Mol Cell, 2008, 30: 108–113.

[43] Kobayashi M, Ohura I, Kawakita K, Yokota N, Fujiwara M, Shimamoto K, Doke N, Yoshioka H. Calcium-dependent protein kinases regulate the production of reactive oxygen species by potato NADPH oxidase. Plant Cell, 2007, 19: 1065–1080.

[44] Suzuki Y, Ishiyama K, Sugawara M, Suzuki Y, Kondo E, Takegahara-Tamakawa Y, Yoon D K, Suganami M, Wada S, Miyake C, et al. Overproduction of chloroplast glyceraldehyde-3-phosphate dehydrogenase improves photosynthesis slightly under elevated [CO₂] conditions in rice. Plant Cell Physiol, 2021, 62: 156–165.

[45] 李敏, 伍国强, 魏明, 刘晨. 植物CDPK在响应逆境胁迫中的作用及机制. 生物工程学报, 2024, 40: 3337–3359.
Li M, Wu G Q, Wei M, Liu C. Functions and mechanisms of CDPKs in plant responses to abiotic stress. Chin J Biotechnol, 2024, 40: 3337–3359 (in Chinese with English abstract).

[46] Zou J J, Li X D, Ratnasekera D, Wang C, Liu W X, Song L F, Zhang W Z, Wu W H. Arabidopsis CALCIUM-DEPENDENT PROTEIN KINASE8 and CATALASE3 function in abscisic acid-mediated signaling and H₂O₂ homeostasis in stomatal guard cells under drought stress. Plant Cell, 2015, 27: 1445–1460.

[47] Zhu C G, Jing B Y, Lin T, Li X Y, Zhang M, Zhou Y H, Yu J Q, Hu Z J. Phosphorylation of sugar transporter TST2 by protein kinase CPK27 enhances drought tolerance in tomato. Plant Physiol, 2024, 195: 1005–1024.