[1] Li X, Wu Y, Zhang D Z, Gallikin J W, Franceschi V R, Okita T W.Rice prolamine protein body biogenesis: a BiP mediated process.Science, 1993, 242: 1054–1056

[2] Okita T W, Li X, Roberts M W. Targeting of mRNAs to domains of the endoplasmic reticulum. Trends Cell Biol, 1994, 4: 91–96

[3] Choi S B, Wang C, Muench D G, Ozawa K, Franceschi V R, Wu Y, Okita T W. Messenger RNA targeting of rice seed storage proteins to specific ER subdomains. Nature, 2000, 407: 765–767

[4] Coleman C E, Lopes M A, Gillikei J W, Boston R S, Larkins B A.A defective signal peptide in the maize high-lysine mutant fleury-2. Proc Natl Acad Sic USA, 1995, 92: 6828–6831

[5] Boehm R, Susanne S, Klaus S, Li S M, Lutz H D. Active expression of the ubiA gene from E. coli in tobacco: influence of plant ER-specific signal peptides on the expression of a membranebound prenyltransferase in plant cells. Transgenic Res, 2000, 9:477–486

[6] Yamagata H, Sugimoto T, Tanaka K, Kasai Z. Biosynthesis of storage proteins in developing rice seeds. Plant Physiol, 1982, 70: 1094–1100

[7] 刘巧泉, 周丽慧, 王红梅, 顾铭洪. 水稻种子贮藏蛋白合成的分子生物学研究进展. 分子植物育种, 2008, 6: 1–15

Liu Q Q, Zhou L H, Wang H M, Gu M H. Advances on biosynthesis of rice seed storage proteins in molecular biology. Mol Plant Breed, 2008, 6: 1–15 (in Chinese with English abstract)

[8] Ren Y L, Wang Y H, Liu F, Zhou K N, Ding Y, Zhou F, Wang Y,Liu K, Gan L, Ma W W, Han X H, Zhang X, Guo X P, Wu F Q,Cheng Z J, Wang J L, Lei C L, Lin Q B, Jiang L, Wu C Y, Bao Y Q, Wang H Y, Wan J M. GLUTELIN PRECURSOR ACCUMULATION3 encodes a regulator of post-Golgi vesicular traffic essential for vacuolar protein sorting in rice endosperm.

Plant Cell, 2014, 26: 410–425

[9] Zhao W M, Gatehouse J A, Boulter D. The purification and partial amino acid sequence of a polypeptide from the glutelin fraction of rice grains: homology to pea legumin. FEBS Lett, 1983,162: 96–102

[10] Wen T N, Luthe D S. Biochemical characterization of rice glutelin. Plant Physiol, 1985, 78: 172–177

[11] Takaiwa F, Oono K, Wing D, Kato A. Sequence of three members and expression of a new major subfamily of glutelin genes from rice. Plant Mol Biol, 1991, 17: 875–885

[12] Okita T W, Hwang Y S, Hnilo J, Kim W T, Aryan A P, Larson R, Krishnan H B. Structure and expression of the rice glutelin multigene family. J Biol Chem, 1989, 264: 12573–12581

[13] Takaiwa F, Ebinuma H, Kikuchi S, Oono K. Nucleotide sequence of a rice glutelin gene. FEBS Lett, 1987, 221: 43–47

[14] Takaiwa F, Kikuchi S, Oono K. A rice glutelin family: a major type of glutelin mRNAs can be divided into two classes. Mol Gen Genet, 1987, 208: 15–22

[15] Takaiwa F, Oono K. Genomic DNA sequences of two new genes for new storage protein glutelin in rice. Jpn J Genet, 1991, 66:161–171

[16] 刘巧泉, 于恒秀, 张文娟, 龚志云, 顾铭洪. 番茄rbcS3A 启动子控制的外源基因在转基因水稻中的表达特性. 植物生理与分子生物学学报, 2007, 33: 251–257

Liu Q Q, Yu H X, Zhang W J, Gong Z Y, Gu M H. Expression of the GUS fusion gene controlled by the tomato rbcS3A promoter in transgenic rice. J Plant Physiol Mol Biol, 2007, 33: 251–257(in Chinese with English abstract)

[17] 刘巧泉, 张景六, 王宗阳, 洪孟民, 顾铭洪. 根癌农杆菌介导的水稻高效转化系统的建立, 植物生理学报, 1998, 24:259–271

Liu Q Q, Zhang J L, Wang Z Y, Hong M M, Gu M H. A highly efficient transformation mediated by Agrobacterium in rice. Acta Phytophysiol Sin, 1998, 24: 259–271 (in Chinese with English abstract)

[18] Murray M G, Thompson W F. Rapid isolation of high molecular weight plant DNA. Nucl Acid Res, 1980, 8: 4321–4325

[19] 郑霏琴, 王宗阳, 高继平. 水稻胚乳中核糖核酸的分离. 植物生理学通讯, 1993, 29: 438–440

Zheng F Q, Wang Z Y, Gao J P. Isolation of nucleic acids from rice endosperm. Plant Physiol Commun, 1993, 29: 438–440 (inChinese with English abstract)

[20] Sambrook J, Fritsch E F, Maniatis T. Molecular Cloning: a Laboratory Manual. Cold Spring Harbor, NY: Cold Spring Harbor Laboratory Press, 1989

[21] Bradford H M. A rapid and sensitive method for the quantification of microgram quantities of protein utilizing the principle of protein-dye binding. Anal Biochem, 1976, 72: 248–254

[22] Jefferson R A. Assaying chimeric genes in plants: The GUS gene fusion system. Plant Mol Biol Rep, 1987, 5: 387–405

[23] Fukuda M, Wen L, Satoh-Cruz M, Kawagoe Y, Nagamura Y,Okita T W, Washida H, Sugino A, Ishino S, Ishino Y, Ogawa M,Sunada M, Ueda T, Kumamaru T. A guanine nucleotide exchange factor for Rab5 proteins is essential for intracellular transport of the proglutelin from the Golgi apparatus to the protein storage vacuole in rice endosperm. Plant Physiol, 2013, 162: 663–674

[24] Mitsukawa N, Konishi R, Uchiki M, Masumura T, Tanaka K. Molecular cloning and characterization of a cysteine-rich16.6-kDa prolamin in rice seeds. Biosci Biotechnol Biochem,

1999, 63: 1851–1858

[25] Boehm R, Susanne S, Klaus S, Li S M, Lutz H D. Active expression of the ubiA gene from E. coli in tobacco: influence of plant ER-specific signal peptides on the expression of a membrane- bound prenyltransferase in plant cells. Transgenic Res,2000, 9: 477–486

[26] Wright K E, Prior F, Sardana R, Altosaar I, Dudani A K, Ganz P R, Tackaberry E S. Sorting of glycoprotein B from human cytomegalovirus to protein storage vesicles in seeds of transgenic tobacco. Transgenic Res, 2001, 10: 177–181

[27] 范云六, 张春义. 迎接21世纪农作物生物技术的挑战, 生物技术通报, 1999, (5): 1–6

Fan Y L, Zhang C Y. Greeting the challenges of crop biotechnology in the 21st century. Biotechnol Inf, 1999, (5): 1–6 (in Chinese with English abstract)

[28] Ye X D, Al-Babili S, Klöti A, Zhang J, Lucca P, Beyer P, Potrykus I. Engineering the provitamin A (beta-carotene) biosynthetic pathway into (carotenoid-free) rice endosperm. Science,

2000, 287: 303–305

[29] Sun S S M, Liu Q Q. Transgenic approaches to improve the nutritional quality of plant proteins. In Vitro Cell Dev Biol-Plant, 2004, 40: 155–162

[30] Fischer R, Stoger E, Schillberg S, Christou P, Twyman R M. Plant-based production of biopharmaceuticals. Curr Opin Plant Biol, 2004, 7: 152–158

[31] Rybicki E P. Plant-made vaccines for humans and animals. Plant Biotech J, 2010, 8: 620–637

[32] Yang Z Q, Liu Q Q, Pan Z M, Yu H X, Jiao X A. Expression of the fusion glycoprotein of newcasstle disease virus in transgenic rice and its immunogenicity in mice. Vaccine, 2007, 25: 591–598

[33] Cheung S C K, Liu L Z, Sun S S M, Liu Q Q, Lan L L, Chan J,Tong P. Inhibition of human MCF-7 breast cancer cells and HT-29 colon cancer cells by rice-produced recombinant human insulin-like growth binding protein-3 (rhIGFBP-3). PLoS One,2013, 8: e77516

[34] Bundó M, Montesinos L, Izquierdo E, Campo S, Mieulet D,Guiderdoni E, Rossignol M, Badosa E, Montesinos E, San Segundo B, Coca M. Production of cecropin A antimicrobial

peptide in rice seed endosperm. BMC Plant Biol, 2014, 14:102